Steven E. Churchill | Scholars@Duke

Steven E. Churchill
Professor in the Department of Evolutionary Anthropology

I am a human paleontologist studying morphological and behavioral adaptation in archaic and modern humans of the Middle and Late Pleistocene. Through comparative functional-morphological analysis of human fossil remains, coupled with investigation of the archeological record of prehistoric human behavior, my students and I conduct research in the following inter-related areas:

1) The ecology, energetics and adaptive strategies of premodern members of the genus Homo (especially the Neandertals [Homo neanderthalensis] of Europe and western Asia and Middle Pleistocene archaic humans of Africa [variously attributed to H. heidelbergensis, H. rhodesiensis or H. helmei] ) and early members of our own species [H. sapiens] in Africa, the Near East and Europe.

2) The evolution of human subsistence strategies across the Middle and Late Pleistocene, with an emphasis on the nature of the hunting methods employed by various groups.

3) The evolution of subsistence technology, especially the origins of true long-range projectile weaponry.

4) The community ecology of humans and large-bodied carnivores in Pleistocene Europe and Africa.

In addition to this basic research, our team is also actively engaged in fieldwork in southern Africa, with the goal of improving our understanding of the morphology and behavior of Middle Stone Age-associated early modern humans and their immediate ancestors (African Middle Pleistocene archaic humans).

Current Appointments & Affiliations

Professor in the Department of Evolutionary Anthropology, Evolutionary Anthropology, Trinity College of Arts & Sciences 2013

Contact Information

Duke Box 90383, Durham, NC 27708-0383
Duke University, Evolutionary Anthropology, 130 Science Drive, Room 108, Durham, NC 27708
churchy@duke.edu (919) 660-7314
http://www.baa.duke.edu/FacPages/churchil.html

Background
Education, Training, & Certifications
- Ph.D., University of New Mexico 1994
- M.A., University of New Mexico 1989
- B.S., Virginia Polytech Institute and State University 1981
Duke Appointment History
- Associate Professor of Biological Anthropology and Anatomy, Evolutionary Anthropology, Trinity College of Arts & Sciences 2002 - 2013
- Chair, Department of Biological Anthropology and Anatomy, Basic Science Departments, School of Medicine 2004 - 2007
- Assistant Professor of Biological Anthropology and Anatomy, Basic Science Departments, School of Medicine 1999 - 2002
- Assistant Professor of Biological Anthropology and Anatomy, Basic Science Departments, School of Medicine 1995 - 1999
Recognition
In the News
- FEB 22, 2016 CBS News
  
  Download and 3D-print real fossils for free
- JAN 5, 2016
  
  Using Digital Tools on Stone Age Treasures
- SEP 17, 2015
  
  Men's Basketball Visits the White House; Stargazing: The Week at Duke {in 60 Seconds}
- SEP 11, 2015 “NBC Nightly News"
  
  Steven Churchill : ‘My mind is absolutely blown’ over human ancestor discovery
- SEP 11, 2015 CNN
  
  Steven Churchill comments: Homo naledi is related to me how?
- SEP 10, 2015 National Geographic
  
  Steve Churchill comments: This face changes the human story
- AUG 4, 2014 WFTV-Orlando, Newsy
  
  Ancient skulls show civilization rose as testosterone fell
- AUG 4, 2014 Daily Telegraph
  
  Humans started making art when their personalities got 'gentler'
- AUG 1, 2014
  
  Society Bloomed With Gentler Personalities and More Feminine Faces
- MAR 6, 2014 Duke Research Blog
  
  New Fossil Cache Hints at Human Ancestry
Expertise
Global Scholarship
- Research
  - Africa (Continent)
  - Europe (Continent)
Research
Selected Grants
- Dissertation Research: Determinants of Form in the Pubis of Middle-to-Late Pleistocene Homo awarded by National Science Foundation 2001 - 2004
- Cultures in Contact: The Italian-American Riparo Bombrini Project awarded by Josiah Charles Trent Memorial Foundation 2001 - 2002
Publications & Artistic Works
Selected Publications
- Books
  - De Ruiter, DJ, Churchill, SE, Hawks, J, and Berger, LR. Late australopiths and the emergence of homo. October 23, 2017. Full Text
  - Churchill, SE. Thin on the Ground: Neandertal Biology, Archeology and Ecology. October 6, 2014. Full Text
- Academic Articles
  - Di Vincenzo, F, Churchill, SE, Buzi, C, Profico, A, Tafuri, MA, Micheli, M, Caramelli, D, and Manzi, G. "Distinct among Neanderthals: The scapula of the skeleton from Altamura, Italy." Quaternary Science Reviews (November 2018). Full Text
  - Holt, B, Negrino, F, Riel-Salvatore, J, Formicola, V, Arellano, A, Arobba, D, Boschian, G, Churchill, SE, Cristiani, E, Di Canzio, E, and Vicino, G. "The Middle-Upper Paleolithic transition in Northwest Italy: new evidence from Riparo Bombrini (Balzi Rossi, Liguria, Italy)." Quaternary International (November 2018). Full Text
  - Yapuncich, GS, Churchill, SE, Cameron, N, and Walker, CS. "Morphometric panel regression equations for predicting body mass in immature humans." American Journal of Physical Anthropology 166, no. 1 (May 2018): 179-195. Full Text
  - Walker, CS, Yapuncich, GS, Sridhar, S, Cameron, N, and Churchill, SE. "Evaluating morphometric body mass prediction equations with a juvenile human test sample: accuracy and applicability to small-bodied hominins." Journal of Human Evolution 115 (February 2018): 65-77. Full Text
  - VanSickle, C, Cofran, Z, García-Martínez, D, Williams, SA, Churchill, SE, Berger, LR, and Hawks, J. "Homo naledi pelvic remains from the Dinaledi Chamber, South Africa." Journal of human evolution (November 20, 2017). Full Text
  - Garvin, HM, Elliott, MC, Delezene, LK, Hawks, J, Churchill, SE, Berger, LR, and Holliday, TW. "Body size, brain size, and sexual dimorphism in Homo naledi from the Dinaledi Chamber." Journal of human evolution 111 (October 2017): 119-138. Full Text
  - Hawks, J, Elliott, M, Schmid, P, Churchill, SE, Ruiter, DJD, Roberts, EM, Hilbert-Wolf, H, Garvin, HM, Williams, SA, Delezene, LK, Feuerriegel, EM, Randolph-Quinney, P, Kivell, TL, Laird, MF, Tawane, G, DeSilva, JM, Bailey, SE, Brophy, JK, Meyer, MR, Skinner, MM, Tocheri, MW, VanSickle, C, Walker, CS, Campbell, TL, Kuhn, B, Kruger, A, Tucker, S, Gurtov, A, Hlophe, N, Hunter, R, Morris, H, Peixotto, B, Ramalepa, M, Rooyen, DV, Tsikoane, M, Boshoff, P, Dirks, PH, and Berger, LR. "New fossil remains of Homo naledi from the Lesedi Chamber, South Africa." Elife 6 (May 9, 2017). Full Text
  - Churchill, SE, and Vansickle, C. "Pelvic Morphology in Homo erectus and Early Homo." Anatomical record (Hoboken, N.J. : 2007) 300, no. 5 (May 2017): 964-977. Full Text
  - Feuerriegel, EM, Green, DJ, Walker, CS, Schmid, P, Hawks, J, Berger, LR, and Churchill, SE. "The upper limb of Homo naledi." Journal of Human Evolution 104 (March 2017): 155-173. Full Text
  - Marchi, D, Walker, CS, Wei, P, Holliday, TW, Churchill, SE, Berger, LR, and DeSilva, JM. "The thigh and leg of Homo naledi." Journal of human evolution 104 (March 2017): 174-204. Full Text
  - Meyer, MR, Williams, SA, Schmid, P, Churchill, SE, and Berger, LR. "The cervical spine of Australopithecus sediba." Journal of human evolution 104 (March 2017): 32-49. Full Text
  - Williams, SA, García-Martínez, D, Bastir, M, Meyer, MR, Nalla, S, Hawks, J, Schmid, P, Churchill, SE, and Berger, LR. "The vertebrae and ribs of Homo naledi." Journal of human evolution 104 (March 2017): 136-154. Full Text
  - Randolph-Quinney, PS, Williams, SA, Steyn, M, Meyer, MR, Smilg, JS, Churchill, SE, Odes, EJ, Augustine, T, Tafforeau, P, and Berger, LR. "Osteogenic tumour in Australopithecus sediba: Earliest hominin evidence for neoplastic disease." South African Journal of Science 112, no. 7-8 (July 1, 2016). Full Text
  - Churchill, SE, Walker, CS, and Schwartz, AM. "Home-range size in large-bodied carnivores as a model for predicting neandertal territory size." Evolutionary anthropology 25, no. 3 (May 2016): 117-123. Full Text
  - Kivell, TL, Deane, AS, Tocheri, MW, Orr, CM, Schmid, P, Hawks, J, Berger, LR, and Churchill, SE. "The hand of Homo naledi." Nature Communications 6 (October 6, 2015): 8431-null. Full Text
  - Dirks, PHGM, Berger, LR, Roberts, EM, Kramers, JD, Hawks, J, Randolph-Quinney, PS, Elliott, M, Musiba, CM, Churchill, SE, de Ruiter, DJ, Schmid, P, Backwell, LR, Belyanin, GA, Boshoff, P, Hunter, KL, Feuerriegel, EM, Gurtov, A, Harrison, JDG, Hunter, R, Kruger, A, Morris, H, Makhubela, TV, Peixotto, B, and Tucker, S. "Geological and taphonomic context for the new hominin species Homo naledi from the Dinaledi Chamber, South Africa." eLife 4 (September 10, 2015). Full Text
  - Berger, LR, Hawks, J, de Ruiter, DJ, Churchill, SE, Schmid, P, Delezene, LK, Kivell, TL, Garvin, HM, Williams, SA, DeSilva, JM, Skinner, MM, Musiba, CM, Cameron, N, Holliday, TW, Harcourt-Smith, W, Ackermann, RR, Bastir, M, Bogin, B, Bolter, D, Brophy, J, Cofran, ZD, Congdon, KA, Deane, AS, Dembo, M, Drapeau, M, Elliott, MC, Feuerriegel, EM, Garcia-Martinez, D, Green, DJ, Gurtov, A, Irish, JD, Kruger, A, Laird, MF, Marchi, D, Meyer, MR, Nalla, S, Negash, EW, Orr, CM, Radovcic, D, Schroeder, L, Scott, JE, Throckmorton, Z, Tocheri, MW, VanSickle, C, Walker, CS, Wei, P, and Zipfel, B. "Homo naledi, a new species of the genus Homo from the Dinaledi Chamber, South Africa." Elife 4 (January 2015). Full Text
  - Harcourt-Smith, WEH, Throckmorton, Z, Congdon, KA, Zipfel, B, Deane, AS, Drapeau, MSM, Churchill, SE, Berger, LR, and DeSilva, JM. "The foot of Homo naledi." Nature Communications 6 (January 2015): 8432-null. Full Text
  - Macias, ME, and Churchill, SE. "Functional Morphology of the Neandertal Scapular Glenoid Fossa." ANATOMICAL RECORD-ADVANCES IN INTEGRATIVE ANATOMY AND EVOLUTIONARY BIOLOGY 298, no. 1 (January 2015): 168-179. Full Text Link to Item
  - Ostrofsky, KR, and Churchill, SE. "Sex Determination by Discriminant Function Analysis of Lumbar Vertebrae." JOURNAL OF FORENSIC SCIENCES 60, no. 1 (January 2015): 21-28. Full Text Link to Item
  - Cieri, RL, Churchill, SE, Franciscus, RG, Tan, J, and Hare, B. "Craniofacial feminization, social tolerance, and the origins of behavioral modernity." Current Anthropology 55, no. 4 (January 1, 2014): 419-443. Full Text
  - Schmid, P, Churchill, SE, Nalla, S, Weissen, E, Carlson, KJ, de Ruiter, DJ, and Berger, LR. "Mosaic Morphology in the Thorax of Australopithecus sediba." SCIENCE 340, no. 6129 (April 12, 2013). Full Text Link to Item
  - Williams, SA, Ostrofsky, KR, Frater, N, Churchill, SE, Schmid, P, and Berger, LR. "The Vertebral Column of Australopithecus sediba." SCIENCE 340, no. 6129 (April 12, 2013). Full Text Link to Item
  - de Ruiter, DJ, DeWitt, TJ, Carlson, KB, Brophy, JK, Schroeder, L, Ackermann, RR, Churchill, SE, and Berger, LR. "Mandibular Remains Support Taxonomic Validity of Australopithecus sediba." SCIENCE 340, no. 6129 (April 12, 2013). Full Text Link to Item
  - Churchill, SE, Holliday, TW, Carlson, KJ, Jashashvili, T, Macias, ME, Mathews, S, Sparling, TL, Schmid, P, de Ruiter, DJ, and Berger, LR. "The upper limb of Australopithecus sediba." Science (New York, N.Y.) 340, no. 6129 (April 2013): 1233477-null. Full Text
  - DeSilva, JM, Holt, KG, Churchill, SE, Carlson, KJ, Walker, CS, Zipfel, B, and Berger, LR. "The lower limb and mechanics of walking in Australopithecus sediba." Science (New York, N.Y.) 340, no. 6129 (April 2013): 1232999-null. Full Text
  - Vincenzo, FD, Churchill, SE, and Manzi, G. "The Vindija Neanderthal scapular glenoid fossa: Comparative shape analysis suggests evo-devo changes among Neanderthals." Journal of Human Evolution 62, no. 2 (2012): 274-285. Full Text
  - Pickering, R, Dirks, PHGM, Jinnah, Z, de Ruiter, DJ, Churchill, SE, Herries, AIR, Woodhead, JD, Hellstrom, JC, and Berger, LR. "Australopithecus sediba at 1.977 Ma and Implications for the Origins of the Genus Homo." SCIENCE 333, no. 6048 (September 9, 2011): 1421-1423. Full Text Link to Item
  - Kibii, JM, Churchill, SE, Schmid, P, Carlson, KJ, Reed, ND, de Ruiter, DJ, and Berger, LR. "A partial pelvis of Australopithecus sediba." Science (New York, N.Y.) 333, no. 6048 (September 8, 2011): 1407-1411. Full Text
  - Kivell, TL, Kibii, JM, Churchill, SE, Schmid, P, and Berger, LR. "Australopithecus sediba hand demonstrates mosaic evolution of locomotor and manipulative abilities." Science (New York, N.Y.) 333, no. 6048 (September 8, 2011): 1411-1417. Full Text
  - Zipfel, B, DeSilva, JM, Kidd, RS, Carlson, KJ, Churchill, SE, and Berger, LR. "The foot and ankle of Australopithecus sediba." Science (New York, N.Y.) 333, no. 6048 (September 8, 2011): 1417-1420. Full Text
  - de Ruiter, DJ, Churchill, SE, Brophy, JK, and Berger, LR. "Regional survey for Middle Stone Age deposits of the Virginia-Theunissen area of the Free State." Navorsinge van die Nasionale Museum Bloemfontein 27 (2011): 1-20. (Academic Article)
  - Lewis, PJ, Johnson, E, Buchanan, B, and Churchill, SE. "The impact of changing grasslands on Late Quaternary bison of the Southern Plains." Quaternary International 217, no. 1-2 (April 15, 2010): 117-130. Full Text
  - Berger, LR, de Ruiter, DJ, Churchill, SE, Schmid, P, Carlson, KJ, Dirks, PHGM, and Kibii, JM. "Australopithecus sediba: a new species of Homo-like australopith from South Africa." Science (New York, N.Y.) 328, no. 5975 (April 2010): 195-204. Full Text
  - Dirks, PHGM, Kibii, JM, Kuhn, BF, Steininger, C, Churchill, SE, Kramers, JD, Pickering, R, Farber, DL, Mériaux, A-S, Herries, AIR, King, GCP, and Berger, LR. "Geological setting and age of Australopithecus sediba from southern Africa." Science (New York, N.Y.) 328, no. 5975 (April 2010): 205-208. Full Text
  - Hartstone-Rose, A, Werdelin, L, De Ruiter, DJ, Berger, LR, and Churchill, SE. "The Plio-Pleistocene ancestor of wild dogs, Lycaon sekowei n. sp." Journal of Paleontology 84, no. 2 (March 1, 2010): 299-308. Full Text
  - Villotte, S, Churchill, SE, Dutour, OJ, and Henry-Gambier, D. "Subsistence activities and the sexual division of labor in the European Upper Paleolithic and Mesolithic: Evidence from upper limb enthesopathies." Journal of Human Evolution 59, no. 1 (2010): 35-43. Full Text
  - Churchill, SE, Franciscus, RG, McKean-Peraza, HA, Daniel, JA, and Warren, BR. "Shanidar 3 Neandertal rib puncture wound and paleolithic weaponry." J Hum Evol 57, no. 2 (August 2009): 163-178. Full Text Link to Item
  - de Ruiter, DJ, Pickering, R, Steininger, CM, Kramers, JD, Hancox, PJ, Churchill, SE, Berger, LR, and Backwell, L. "New Australopithecus robustus fossils and associated U-Pb dates from Cooper's Cave (Gauteng, South Africa)." Journal of Human Evolution 56, no. 5 (May 13, 2009): 497-513. Full Text
  - Froehle, AW, and Churchill, SE. "Energetic competition between Neandertals and anatomically modern humans." Paleoanthropology (2009): 96-116. (Academic Article)
  - Rhodes, JA, and Churchill, SE. "Throwing in the Middle and Upper Paleolithic: inferences from an analysis of humeral retroversion." Journal of Human Evolution 56, no. 1 (2009): 1-10. Full Text
  - Berger, LR, Churchill, SE, Klerk, BD, and Quinn, RL. "Small-bodied humans from Palau, Micronesia." PLoS ONE 3, no. 3 (2008). Full Text Open Access Copy
  - Ruiter, DJD, Brophy, JK, Lewis, PJ, Churchill, SE, and Berger, LR. "Faunal assemblage composition and paleoenvironment of Plovers Lake, a Middle Stone Age locality in Gauteng Province, South Africa." Journal of Human Evolution 55, no. 6 (2008): 1102-1117. Full Text
  - Hartstone-Rose, A, De Ruiter, DJ, Berger, LR, and Churchill, SE. "A sabre-tooth felid from Coopers Cave (Gauteng, South Africa) and its implications for Megantereon (Felidae: Machairodontinae) taxonomy." Palaeontologia Africana 42 (May 1, 2007): 99-108.
  - Hartstone Rose, A, de Ruiter, DJ, Berger, LR, and Churchill, SE. "A saber-tooth felid from Coopers Cave (Gauteng, South Africa) and its implications for Megantereon taxonomy." Paleontologica Africana 42 (2007): 99-108. (Academic Article)
  - Lewis, PJ, Johnson, E, Buchanan, B, and Churchill, SE. "The evolution of Bison bison: A view from the Southern Plains." Bull. Texas Arch. Soc. 78 (2007): 197-204. (Academic Article)
  - Yokley, TR, and Churchill, SE. "Archaic and modern human distal humeral morphology." J Hum Evol 51, no. 6 (December 2006): 603-616. Full Text Link to Item
  - Churchill, SE, and Rhodes, JA. "How strong were the Neandertals? Leverage and muscularity at the shoulder and elbow in mousterian foragers." Periodicum Biologorum 108, no. 4 (August 1, 2006): 457-470.
  - Brophy, JK, de Ruiter, DJ, Lewis, PJ, Churchill, SE, and Berger, LR. "Preliminary investigation of the new Middle Stone Age site of Plovers Lake, South Africa." Curr. Res. Pleistocene 23 (2006): 41-43. (Academic Article)
  - Churchill, SE, Shackelford, LL, Georgi, JN, and Black, MT. "Morphological variation and airflow dynamics in the human nose." Am J Hum Biol 16, no. 6 (November 2004): 625-638. Full Text Link to Item
  - Brink, , S, J, Berger, LR, Churchill, SE, Rossouw, L, and d, DSN. "Mid-Late Quaternary mammalian fossils from the Sand River drainage, Free State Province, central southern Africa (In preparation)." Paleontol. Afr. (2004). (Academic Article)
  - Schmitt, D, Churchill, SE, and Hylander, WL. "Experimental evidence concerning spear use in Neandertals and early modern humans." Journal of Archaeological Science 30, no. 1 (January 1, 2003): 103-114. Full Text
  - "Of Assegais and Bayonets: Reconstructing Prehistoric Spear Use." Evolutionary Anthropology 11, no. 5 (December 1, 2002): 185-186. (Review) Full Text
  - Franciscus, RG, and Churchill, SE. "The costal skeleton of Shanidar 3 and a reappraisal of Neandertal thoracic morphology." Journal of Human Evolution 42, no. 3 (March 2002): 303-356. Full Text
  - Churchill, SE. "Hand morphology, manipulation, and tool use in Neandertals and early modern humans of the Near East." Proceedings of the National Academy of Sciences of the United States of America 98, no. 6 (March 2001): 2953-2955. (Review) Full Text
  - Churchill, SE, and Smith, FH. "A modern human humerus from the early aurignacian of Vogelherdhöhle (Stetten, Germany)." American Journal of Physical Anthropology 112, no. 2 (June 2000): 251-273. Full Text
  - Churchill, SE, Berger, LR, and Parkington, JE. "A Middle Pleistocene human tibia from Hoedjiespunt, Western Cape, South Africa." South African Journal of Science 96, no. 7 (2000): 367-368.
  - Churchill, SE, Brink, JS, Berger, LR, Hutchison, RA, Rossouw, L, Stynder, D, Hancox, PJ, Brandt, D, Woodborne, S, Loock, JC, Scott, L, and Ungar, P. "Erfkroon: A new Florisian fossil locality from fluvial contexts in the western Free State, South Africa." South African Journal of Science 96, no. 4 (2000): 161-163.
  - Churchill, SE, and Smith, FH. "Makers of the early Aurignacian of Europe." Am J Phys Anthropol Suppl 31 (2000): 61-115. Link to Item
  - Holt, BM, Mussi, M, Churchill, SE, and Formicola, V. "Biological and cultural trends in Upper Palaeolithic Europe." Riv. Anthropol 78 (2000): 179-192. (Academic Article)
  - Trinkaus, E, Churchill, SE, Ruff, CB, and Vandermeersch, B. "Long bone shaft robusticity of body proportions of the Saint-Cesaire 1 and Chatelperronian Neanderthal." Journal of Archaeological Science 26, no. 7 (January 1, 1999): 753-773. Full Text
  - Trinkaus, E, and Churchill, SE. "Diaphyseal cross-sectional geometry of Near Eastern Middle Palaeolithic humans: The humerus." Journal of Archaeological Science 26, no. 2 (January 1, 1999): 173-184. Full Text
  - Hamrick, MW, Churchill, SE, Schmitt, D, and Hylander, WL. "EMG of the human flexor pollicis longus muscle: implications for the evolution of hominid tool use." Journal of Human Evolution 34, no. 2 (February 1998): 123-136. Full Text
  - Churchill, SE, and Morris, AG. "Muscle Marking Morphology and Labour Intensity in Prehistoric Khoisan Foragers." International Journal of Osteoarchaeology 8, no. 5 (January 1, 1998): 390-411. Full Text
  - Churchill, SE. "Cold adaptation, heterochrony, and neandertals." Evolutionary Anthropology: Issues, News, and Reviews 7, no. 2 (1998): 46-60. Full Text
  - Pearson, OM, Churchill, SE, Grine, FE, Trinkaus, E, and Holliday, TW. "Multivariate analyses of the hominid ulna from Klasies River Mouth." Journal of Human Evolution 34, no. 6 (1998): 653-656.
  - Trinkaus, E, Ruff, CB, Churchill, SE, and Vandermeersch, B. "Locomotion and body proportions of the Saint-Cesaire 1 Chatelperronian Neandertal." Proceedings of the National Academy of Sciences of the United States of America 95, no. 10 (1998): 5836-5840. Full Text
  - Byers, SN, Churchill, SE, and Curran, B. "Identification of Euro-Americans, Afro-Americans, and Amerindians from palatal dimensions." Journal of Forensic Sciences 42, no. 1 (1997): 3-9.
  - Churchill, SE, and Formicola, V. "A Case of Marked Bilateral Asymmetry in the Upper Limbs of an Upper Palaeolithic Male from Barma Grande (Liguria), Italy." International Journal of Osteoarchaeology 7, no. 1 (January 1, 1997): 18-38. Full Text
  - Churchill, SE, Pearson, OM, Grine, FE, Trinkaus, E, and Holliday, TW. "Morphological affinities of the proximal ulna from Klasies River main site: archaic or modern?." Journal of Human Evolution 31, no. 3 (September 1996): 213-237. Full Text
  - Churchill, SE. "Particulate versus integrated evolution of the upper body in late pleistocene humans: a test of two models." American Journal of Physical Anthropology 100, no. 4 (August 1996): 559-583. Full Text
  - Trinkaus, E, Churchill, SE, and Ruff, CB. "Postcranial robusticity in Homo. II: Humeral bilateral asymmetry and bone plasticity." American Journal of Physical Anthropology 93, no. 1 (January 1994): 1-34. Full Text
  - Churchill, SE. "Weapon Technology, Prey Size Selection, and Hunting Methods in Modern Hunter‐Gatherers: Implications for Hunting in the Palaeolithic and Mesolithic." Archeological Papers of the American Anthropological Association 4, no. 1 (January 1, 1993): 11-24. Full Text
  - Trinkaus, E, Churchill, SE, Villemeur, I, Riley, KG, Heller, JA, and Ruff, CB. "Robusticity versus shape: the functional interpretation of Neandertal appendicular morphology." J. Anth. Soc. Nippon 99 (1991): 257-278. (Academic Article)
  - Churchill, SE, and Trinkaus, E. "Neandertal scapular glenoid morphology." American Journal of Physical Anthropology 83, no. 2 (1990): 147-160.
  - Trinkhaus, E, and Churchill, SE. "Neandertal radial tuberosity orientation." American Journal of Physical Anthropology 75, no. 1 (1988): 15-21.
  - Churchill, SE, and Smith, FH. "Makers of the early Aurignacian of Europe." American Journal of Physical Anthropology Suppl 31 (January 1, 1969): 61-115.
- Book Sections
  - Salem, PE, and Churchill, SE. "Penetration, Tissue Damage, and Lethality of Wood- Versus Lithic-Tipped Projectiles." In Vertebrate Paleobiology and Paleoanthropology, 203-212.: Springer Netherlands, 2016. Full Text
  - Walker, CS, and Churchill, SE. "Territory size in Canis lupus: Implications for Neandertal mobility." In Reconstructing Mobility: Environmental, Behavioral, and Morphological Determinants, 209-226. April 1, 2014. Full Text
  - de Ruiter, DJ, Churchill, SE, and Berger, LR. "Australopithecus sediba from Malapa, South Africa." In The Paleobiology of Australopithecus, 147-160.: Springer Netherlands, 2013. Full Text
  - Churchill, SE, and Rhodes, JA. "The Evolution of the Human Capacity for “Killing at a Distance”: The Human Fossil Evidence for the Evolution of Projectile Weaponry." In Vertebrate Paleobiology and Paleoanthropology, 201-210. January 1, 2009. Full Text
  - Churchill, SE. "Bioenergetic perspectives on Neanderthal thermoregulatory and activity budgets." In Neanderthals Revisited: New Approaches and Perspectives, 113-133.: Springer Netherlands, 2006. Full Text
- Conference Papers
  - Cook, RW, Yapuncich, GS, Thompson, IJ, Walker, CS, and Churchill, SE. "A comparison of lateral iliac flare measurement methods and their correlation with lesser gluteal moment arms." 87th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA). Austin, TX. April 11, 2018 - April 14, 2018.: WILEY, April 1, 2018. Link to Item
  - De Ruiter, DJ, Brophy, JK, Van der Merwe, R, Smilg, JS, Churchill, SE, and Berger, LR. "New craniodental remains of the type specimen of Australopithecus sediba." 87th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA). Austin, TX. April 11, 2018 - April 14, 2018.: WILEY, April 1, 2018. Link to Item
  - Feuerriegel, EM, Voisin, J-L, Churchill, SE, Hawks, J, and Berger, LR. "The upper limb of Homo naledi: New material from the Lesedi Chamber, Rising Star System, South Africa." 87th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA). Austin, TX. April 11, 2018 - April 14, 2018.: WILEY, April 1, 2018. Link to Item
  - Friedl, L, Claxton, AG, Walker, CS, Churchill, SE, Holliday, TW, Hawks, J, Berger, LR, Desilva, JM, and Marchi, D. "Femoral neck and shaft structure in Homo naledi." 87th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA). Austin, TX. April 11, 2018 - April 14, 2018.: WILEY, April 1, 2018. Link to Item
  - Green, DJ, Churchill, SE, Macias, ME, Gunz, P, Carlson, KJ, Schmid, P, and Berger, LR. "Three-dimensional morphology and comparative anatomy of the Australopithecus sediba scapula." 87th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA). Austin, TX. April 11, 2018 - April 14, 2018.: WILEY, April 1, 2018. Link to Item
  - Walker, CS, Yapuncich, GS, Bowie, A, Belais, R, and Churchill, SE. "Accuracy of human-based morphometric equations for predicting bonobo body mass." 87th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA). Austin, TX. April 11, 2018 - April 14, 2018.: WILEY, April 1, 2018. Link to Item
  - Williams, SA, Prang, TC, Grabowski, MW, Meyer, MR, Schmid, P, Churchill, SE, and Berger, LR. "Relative size and scaling of the lumbo-sacral joint in fossil hominins: Implications for function and phylogeny." 87th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA). Austin, TX. April 11, 2018 - April 14, 2018.: WILEY, April 1, 2018. Link to Item
  - Bastir, M, Garcia-Martinez, D, Williams, SA, Meyer, MR, Nalla, S, Schmid, P, Barash, A, Oishi, M, Ogihara, N, Churchill, SE, Hawks, J, and Berger, LR. "Geometric morphometrics of hominoid thoraces and its bearing for reconstructing the ribcage of H. naledi." 86th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA). New Orleans, LA. April 19, 2017 - April 22, 2017.: WILEY, April 1, 2017. Link to Item
  - Churchill, SE. "The functional significance of iliac buttressing in the genus Homo." 86th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA). New Orleans, LA. April 19, 2017 - April 22, 2017.: WILEY, April 1, 2017. Link to Item
  - Elliott, MC, Dirks, PHGM, Berger, LR, Roberts, EM, Kramers, JD, Hawks, J, Randolph-Quinney, PS, Musiba, CM, Churchill, SE, de Ruiter, DJ, Schmid, P, Backwell, LR, Belyanin, GA, Boshoff, P, Hunter, KL, Feuerriegel, EM, Gurtov, A, Harrison, JDG, Hunter, R, Kruger, A, Morris, H, and Peixotto, B. "Geological and taphonomic context of excavations within the Rising Star cave system." March 2016. Link to Item
  - Feuerriegel, EM, Green, DJ, Walker, CS, Schmid, P, Hawks, J, Berger, LR, and Churchill, SE. "The shoulder and upper limb of Homo naledi." March 2016. Link to Item
  - Garvin, HM, Elliott, MC, Delezene, LK, Hawks, JD, Churchill, SE, Berger, LR, and Holliday, TW. "Body size and sexual dimorphism in H. naledi." March 2016. Link to Item
  - Kivell, TL, Deane, AS, Tocheri, MW, Orr, CM, Schmid, P, Hawks, J, Berger, LR, and Churchill, SE. "Functional interpretation of the Homo naledi hand." March 2016. Link to Item
  - Marchi, D, Walker, CS, Wei, P, Holliday, TW, Churchill, SE, Berger, LR, and Desilva, JM. "Thigh and leg remains of Homo naledi." March 2016. Link to Item
  - Vansickle, C, Cofran, ZD, Garcia-Martinez, D, Williams, SA, Churchill, SE, Berger, LR, and Hawks, J. "Primitive pelvic features in a new species of Homo." March 2016. Link to Item
  - Walker, CS, Desilva, JM, Holliday, TW, Marchi, D, Garvin, HM, Cofran, Z, Hawks, J, Berger, LR, and Churchill, SE. "Relative length of the immature Homo naledi tibia UW 101-1070: evidence for elongation of the leg." March 2016. Link to Item
  - Williams, SA, Garcia-Martinez, D, Meyer, MR, Nalla, S, Schmid, P, Hawks, J, Churchill, SE, Berger, LR, and Bastir, M. "The axial skeleton and scaling of the trunk in Homo naledi." March 2016. Link to Item
  - Brophy, JK, de Ruiter, DJ, Berger, LR, Churchill, SE, and Schmid, P. "Morphometric analyses of maxillary and mandibular first molars of Pleistocene hominins." March 2015. Link to Item
  - Churchill, SE, Sridhar, S, Cameron, N, and Walker, CS. "Geometric methods of body mass estimation in small-bodied hominins." March 2015. Link to Item
  - Sylvester, AD, Desilva, JM, Churchill, SE, and Berger, LR. "Three-dimensional shape analysis of the distal femur of Australopithecus sediba." March 2015. Link to Item
  - Ostrofsky, KR, Williams, SA, Churchill, SE, Berger, LR, and Richmond, BG. "Australopith lumbar vertebral morphology: Insights from Australopithecus sediba." March 2014. Link to Item
  - Reed, ND, Hakki, L, and Churchill, SE. "Pelvic sexual dimorphism in the hominin fossil record." March 2014. Link to Item
  - Macias, ME, and Churchill, SE. "Functional morphology of the Neandertal scapular glenoid fossa." 2013. Link to Item
  - Ostrofsky, KR, and Churchill, SE. "Sex determination by discriminant function analysis of lumbar vertebrae." 2013. Link to Item
  - Williams, SA, Churchill, SE, Ostrofsky, KR, Schmid, P, Frater, N, and Berger, LR. "The number of vertebrae in early hominins: insights from Australopithecus sediba." 2013. Link to Item
  - Desilva, JM, Zipfel, B, Kidd, RS, Carlson, KJ, Churchill, SE, and Berger, LR. "The primitive aspects of the foot and ankle of Australopithecus sediba." 2012. Link to Item
  - Macias, ME, Wall, CE, and Churchill, SE. "Size and shape in the primate forelimb." 2012. Link to Item
  - Reed, ND, and Churchill, SE. "Acetabulocristal buttressing in hominins." 2012. Link to Item
  - Walker, CS, Mayer, E, and Churchill, SE. "Morphological correlates of human hip osteoarthritis." 2012. Link to Item
  - Yokley, TR, Froehle, AW, and Churchill, SE. "Decreased maintenance energy expenditure in modern human and the resultant demographic displacement of archaic humans." 2012. Link to Item
  - Kivell, TL, Kibii, JM, Churchill, SE, and Berger, LR. "Evolution of the hominin hand: old and new evidence from the Plio-Pleistocene." 2011. Link to Item
  - Walker, CS, and Churchill, SE. "Territory size in Canis lupus: implications for Neandertal mobility." 2011. Link to Item
  - de Ruiter, DJ, Carlson, K, Brophy, J, Carlson, KJ, Churchill, SE, Schmid, P, Kibii, JM, and Berger, LR. "Craniodental remains of Australopithecus sediba from Malapa, South Africa." 2011. Link to Item
  - Hatala, KG, Churchill, SE, Ullinger, J, and Sheridan, SG. "Activity-related changes in the geometry, of the proximal femur: A study of two Near Eastern samples." 2010. Link to Item
  - Brophy, JK, de Ruiter, DJ, Churchill, SE, and Berger, LR. "Accumulating agent and paleoenvironment of the hominin-bearing site of Plovers Lake, South Africa." 2006. Link to Item
  - Berger, LR, Churchill, SE, and de Ruiter, DJ. "Plover's Lake: A hominin-bearing Middle Stone Age site in the Witwatersrand Area, South africa." 2003. Link to Item
  - Miller, SF, Yokley, TR, Churchill, SE, Franciscus, RG, Hublin, JJ, and Eaves-Johnson, KL. "A new technique for reconstructing the vocal anatomy of fossil humans." 2003. Link to Item
  - Yokley, TR, and Churchill, SE. "Archaic and modern human distal humeral morphology." 2002. Link to Item
  - Churchill, SE, Brink, JS, and Gruss, LT. "Middle Pleistocene to Holocene postcranial gracilization in black wildebeest, Connochaetes gnou, and its implications for understanding diachronic changes in robusticity in the genus Homo." 2000. Link to Item
  - Churchill, SE, Shackelford, LL, Georgi, JN, and Black, MT. "Airflow dynamics in the Neandertal nose." April 1999. Link to Item
  - Churchill, SE, Shackelford, LL, Georgi, JN, and Black, MT. "Morphological variation in the upper respiratory tract and airflow dynamics." 1999. Link to Item
  - Georgi, JN, Churchill, SE, Shackelford, LL, and Black, MT. "Nasal valve function and location as determined by in vitro fluid flow studies." 1999. Link to Item
  - Hambrick, MW, Churchill, SE, Schmitt, D, and Hylander, WL. "Tool use in Homo and Australopithecus." April 1997. Link to Item
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- Editorial Advisory board, Human Evolution Series, Oxford University Press. 2008 2008
- Editorial board, Museum of Comparative Anthropogeny. 2008 2008

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