Ram Oren | Scholars@Duke

Ram Oren
Nicholas Professor of Earth Systems Science

With his graduate students, Dr. Oren quantifies the components of water flux in forest ecosystems and the influence of biotic and abiotic factors on water and, due to strong links between carbon and water, on carbon flux. Climate variability, including variations in air temperature, vapor pressure deficit, incoming radiation and soil moisture, and environmental change, including elevated atmospheric carbon dioxide, affect the intra- and inter-annual patterns and amounts of water used by forest ecosystems, and its spatial distribution, as well as carbon uptake and sequestration. The variation of water flux influence the temporal and spatial partitioning of incoming radiation between latent and sensible heat. Using a local mass balance approach and detailed measurements of water flux and driving variables in the soil, plants, and the atmosphere, in forests from Brazil to Alaska, Dr. Oren evaluates the likely responses of different forest ecosystems to environmental change.

Current Appointments & Affiliations

Nicholas Professor of Earth Systems Science, Environmental Sciences and Policy, Nicholas School of the Environment 2009
Professor of Ecology, Environmental Sciences and Policy, Nicholas School of the Environment 2002
Professor in the Department of Civil and Environmental Engineering, Civil and Environmental Engineering, Pratt School of Engineering 2015

Contact Information

A246A Lev Sci Res Ctr, Durham, NC 27708
Box 90328, Durham, NC 27708-0328
ramoren@duke.edu (919) 613-8032

Background
Education, Training, & Certifications
- Ph.D., Oregon State University 1984
- M.S., Oregon State University 1980
- B.S., Humboldt State University 1978
Duke Appointment History
- Professor in the Department of Civil and Environmental Engineering, Civil and Environmental Engineering, Pratt School of Engineering 2009 - 2012
- Chair, Division of Environmental Sciences and Policy, Environmental Sciences and Policy, Nicholas School of the Environment 2007 - 2010
- Associate Professor with Tenure, Nicholas School of the Environment, Duke University 1993 - 2002
- Assistant Professor, Nicholas School of the Environment, Duke University 1986 - 1993
Recognition
In the News
- NOV 27, 2018
  
  Duke Faculty Plentiful On ‘Highly Cited’ List
Expertise
Global Scholarship
- Research
Research
Selected Grants
- Constraining the Simultaneous Effects of Elevated CO2, Temperature, and Shifts in Rainfall Patterns on Ecosystme Carbon awarded by Department of Energy 2011 - 2016
- Can Variation in Hydraulic Conductance Explain Isohydric and Anisohydric Stomatal Behavior? awarded by US-Israel Binational Science Foundation 2011 - 2016
- Duke Forest FACE Experiment: Forest-Atmosphere Carbon Transfer and Storage awarded by Department of Energy 2001 - 2015
- How will the Interaction Between Roots and Canopy Physiological Functioning at Night Influence Water and Carbon Fluxes awarded by North Carolina State University 2011 - 2015
- Hydraulic Controls on Carbon Cycling and Water Exchange Rates of Southern Conifers awarded by Department of Agriculture 2011 - 2015
- The Duke Free Air Co2 Enrichement (FACE) Experiment: Woody Roots Sampling awarded by Forest Service 2011 - 2012
- Effect of Nitrogen Fertilization on Carbon Cycling and Sequestration Under Current and Elevated Atmospheric CO2 awarded by Forest Service 2009 - 2012
- De-convolving the Effects of Rising Atmospheric CO2, Solar Dimming, and Afforestation awarded by National Science Foundation 2006 - 2010
- Controls of Net Ecosystem Exchange at an Old Field, a Pine Plantation, and a Hardwood Forest Under Identical Climatic an awarded by Department of Energy 2000 - 2009
- Forest-Atmosphere Carbon Transfer and Storage (FACTS-I): Continuation awarded by Department of Energy 1995 - 2007
- REDUCING UNCERTAINTY IN CURRENT AND PROJECTED CARBON SEQUESTRATION IN SOUTHEASTERN US: Fellowship Supplement awarded by Forest Service 2005 - 2007
- Reducing Uncertainty in Current and Projected Carbon Sequestration in Southeastern US awarded by Forest Service 2003 - 2006
- Reducing Uncertainty in Current and Projected Carbon Sequestration in Southeastern U.S.: Phase VI awarded by Forest Service 2004 - 2005
- Reducing Uncertainty in Current and Projected Carbon Sequestration in Southeastern US, Phase V.2 awarded by Forest Service 2003 - 2004
- Reducing Uncertainty in Current and Projected Carbon Sequestration in Southeastern US awarded by Forest Service 2001 - 2003
- Reducing Uncertainty in Current and Projected Carbon Sequestration in Southeastern US awarded by Department of Agriculture 2000 - 2003
- Reducing uncertainty in current and projected carbon sequestration in southeastern US awarded by Forest Service 2001 - 2003
- Forest-Atmosphere Carbon Transfer and Storage (FACTS-1) awarded by Office of Energy Research 1995 - 2001
- Reducing uncertainty in current and projected carbon sequestration in southeastern US awarded by Forest Service 2000 - 2001
- (98-0832) Responses of a Loblolly Pine Plantation to Elevated Supplies of C02 and Nutrients awarded by Forest Service 1998 - 1999
- (98-0747) Effects of Long Term Increased Water and Nutrient Supply on Drought Response, Water Relations, and Water Balan awarded by Forest Service 1998 - 1999
- (97-0401) Spatial Variability of Carbon Dioxide Fluxes in a Homogeneous Pine Forest awarded by National Institute for Global Environmental Change 1997 - 1999
- (97-0058) Effects of Water and Nutrient Supply on Water Use in Loblolly Pine awarded by Forest Service 1996 - 1998
- (95-0754) Evapotranspiration in Seasonally-Dry Amazonian Ecosystems: Water Budget Responses to Forest Removal and Regrow awarded by National Aeronautics and Space Administration 1995 - 1996
- (93-0749) NASA Graduate Student Fellowship: Evapotranspiration in Seasonally Dry Amazonian Ecosystems awarded by National Aeronautics and Space Administration 1993 - 1996
- (94-0754) Evapotranspiration in Seasonally Dry Amazonian Ecosystems: Water Budget Responses to Forest Removal and Regrow awarded by National Aeronautics and Space Administration 1993 - 1996
- (95-0230) Evapotranspiration in Seasonally Dry Amazonian Ecosystems awarded by National Aeronautics and Space Administration 1993 - 1996
- (95-0316) Carbon Fluxes in a Managed Pine Forest Under Ambient and Elevated CO2 awarded by National Institute for Global Environmental Change 1995 - 1996
- (94-1009) Predictions and Measurements of Oxidants Precursor Emissions from Forested Ecosystems: Scaling from Leaf to Ca awarded by Environmental Protection Agency 1994 - 1995
- (95-0243) Water-use of Managed Southern Pine and Natural Hardwood Forests awarded by Department of Energy 1992 - 1995
- (92-0377) Water-Use of Managed Southern Pine and Natural Hardwood Forests awarded by Department of Energy 1992 - 1993
- (92-0807) Transpiration, Canopy Conductance, and Water Status of Boreal Plant Communities awarded by Jet Propulsion Lab 1992
- (88-0567) Meeting Nutrient Demands From Retranslocation and Soil Sources: Nitrogen and Phosphorous in Loblolly Pine awarded by Forest Service 1988 - 1990
Publications & Artistic Works
Selected Publications
- Academic Articles
  - Yin, C, Palmroth, S, Pang, X, Tang, B, Liu, Q, and Oren, R. "Differential responses of Picea asperata and Betula albosinensis to nitrogen supply imposed by water availability." Tree Physiology 38, no. 11 (November 2018): 1694-1705. Full Text
  - Ward, EJ, Oren, R, Seok Kim, H, Kim, D, Tor-Ngern, P, Ewers, BE, McCarthy, HR, Oishi, AC, Pataki, DE, Palmroth, S, Phillips, NG, and Schäfer, KVR. "Evapotranspiration and water yield of a pine-broadleaf forest are not altered by long-term atmospheric [CO2 ] enrichment under native or enhanced soil fertility." Global Change Biology 24, no. 10 (October 2018): 4841-4856. Full Text
  - Ryan, MG, Oren, R, and Waring, RH. "Fruiting and sink competition." Tree Physiology 38, no. 9 (September 2018): 1261-1266. Full Text
  - Tor-ngern, P, Oren, R, Palmroth, S, Novick, K, Oishi, A, Linder, S, Ottosson-Löfvenius, M, and Näsholm, T. "Water balance of pine forests: Synthesis of new and published results." Agricultural and Forest Meteorology 259 (September 2018): 107-117. Full Text
  - Montaldo, N, and Oren, R. "Changing Seasonal Rainfall Distribution With Climate Directs Contrasting Impacts at Evapotranspiration and Water Yield in the Western Mediterranean Region." Earth'S Future 6, no. 6 (June 2018): 841-856. Full Text
  - Tarvainen, L, Wallin, G, Lim, H, Linder, S, Oren, R, Ottosson Löfvenius, M, Räntfors, M, Tor-Ngern, P, and Marshall, J. "Photosynthetic refixation varies along the stem and reduces CO2 efflux in mature boreal Pinus sylvestris trees." Tree Physiology 38, no. 4 (April 2018): 558-569. Full Text
  - Kim, D, Oren, R, Clark, JS, Palmroth, S, Oishi, AC, McCarthy, HR, Maier, CA, and Johnsen, K. "Dynamics of soil CO2 efflux under varying atmospheric CO2 concentrations reveal dominance of slow processes." Global change biology 23, no. 9 (September 2017): 3501-3512. Full Text
  - Lim, H, Oren, R, Linder, S, From, F, Nordin, A, Fahlvik, N, Lundmark, T, and Näsholm, T. "Annual climate variation modifies nitrogen induced carbon accumulation of Pinus sylvestris forests." Ecological applications. 27, no. 6 (September 2017): 1838-1851. Full Text
  - Way, DA, Stinziano, JR, Berghoff, H, and Oren, R. "How well do growing season dynamics of photosynthetic capacity correlate with leaf biochemistry and climate fluctuations?." Tree physiology 37, no. 7 (July 2017): 879-888. Full Text
  - Metcalfe, DB, Ricciuto, D, Palmroth, S, Campbell, C, Hurry, V, Mao, J, Keel, SG, Linder, S, Shi, X, Näsholm, T, Ohlsson, KEA, Blackburn, M, Thornton, PE, and Oren, R. "Informing climate models with rapid chamber measurements of forest carbon uptake." Global change biology 23, no. 5 (May 2017): 2130-2139. Full Text
  - Franklin, O, Cambui, CA, Gruffman, L, Palmroth, S, Oren, R, and Näsholm, T. "The carbon bonus of organic nitrogen enhances nitrogen use efficiency of plants." Plant, Cell & Environment 40, no. 1 (January 2017): 25-35. Full Text
  - Tor-Ngern, P, Oren, R, Oishi, AC, Uebelherr, JM, Palmroth, S, Tarvainen, L, Ottosson-Löfvenius, M, Linder, S, Domec, J-C, and Näsholm, T. "Ecophysiological variation of transpiration of pine forests: synthesis of new and published results." Ecological Applications : a Publication of the Ecological Society of America 27, no. 1 (January 2017): 118-133. Full Text
  - Hasselquist, NJ, Metcalfe, DB, Inselsbacher, E, Stangl, Z, Oren, R, Näsholm, T, and Högberg, P. "Greater carbon allocation to mycorrhizal fungi reduces tree nitrogen uptake in a boreal forest." Ecology 97, no. 4 (April 2016): 1012-1022.
  - Kim, D, Oren, R, and Qian, SS. "Response to CO2 enrichment of understory vegetation in the shade of forests." Global change biology 22, no. 2 (February 2016): 944-956. Full Text
  - Montaldo, N, and Oren, R. "The way the wind blows matters to ecosystem water use efficiency." Agricultural and Forest Meteorology 217 (February 2016): 1-9. Full Text
  - Oishi, AC, Hawthorne, DA, and Oren, R. "Baseliner: An open-source, interactive tool for processing sap flux data from thermal dissipation probes." Softwarex 5 (2016): 139-143. Full Text
  - Henriksson, N, Tarvainen, L, Lim, H, Tor-Ngern, P, Palmroth, S, Oren, R, Marshall, J, and Näsholm, T. "Stem compression reversibly reduces phloem transport in Pinus sylvestris trees." Tree physiology 35, no. 10 (October 2015): 1075-1085. Full Text
  - Moshelion, M, Halperin, O, Wallach, R, Oren, R, and Way, DA. "Role of aquaporins in determining transpiration and photosynthesis in water-stressed plants: crop water-use efficiency, growth and yield." Plant, cell & environment 38, no. 9 (September 2015): 1785-1793. (Review) Full Text
  - Zhang, Q, Katul, GG, Oren, R, Daly, E, Manzoni, S, and Yang, D. "The hysteresis response of soil CO2 concentration and soil respiration to soil temperature." Journal of Geophysical Research: Biogeosciences 120, no. 8 (August 2015): 1605-1618. Full Text
  - Attia, Z, Domec, J-C, Oren, R, Way, DA, and Moshelion, M. "Growth and physiological responses of isohydric and anisohydric poplars to drought." Journal of Experimental Botany 66, no. 14 (July 2015): 4373-4381. Full Text Open Access Copy
  - Bell, DM, Ward, EJ, Oishi, AC, Oren, R, Flikkema, PG, and Clark, JS. "A state-space modeling approach to estimating canopy conductance and associated uncertainties from sap flux density data." Tree physiology 35, no. 7 (July 2015): 792-802. Full Text
  - Lim, H, Oren, R, Palmroth, S, Tor-ngern, P, Mörling, T, Näsholm, T, Lundmark, T, Helmisaari, H-S, Leppälammi-Kujansuu, J, and Linder, S. "Inter-annual variability of precipitation constrains the production response of boreal Pinus sylvestris to nitrogen fertilization." Forest Ecology and Management 348 (July 2015): 31-45. Full Text Open Access Copy
  - Medlyn, BE, Zaehle, S, De Kauwe, MG, Walker, AP, Dietze, MC, Hanson, PJ, Hickler, T, Jain, AK, Luo, Y, Parton, W, Prentice, IC, Thornton, PE, Wang, S, Wang, Y-P, Weng, E, Iversen, CM, McCarthy, HR, Warren, JM, Oren, R, and Norby, RJ. "Using ecosystem experiments to improve vegetation models." Nature Climate Change 5, no. 6 (June 2015): 528-534. Full Text
  - Way, DA, Oren, R, and Kroner, Y. "The space-time continuum: the effects of elevated CO2 and temperature on trees and the importance of scaling." Plant, cell & environment 38, no. 6 (June 2015): 991-1007. (Review) Full Text
  - Xiong, W, Oren, R, Wang, Y, Yu, P, Liu, H, Cao, G, Xu, L, Wang, Y, and Zuo, H. "Heterogeneity of competition at decameter scale: patches of high canopy leaf area in a shade-intolerant larch stand transpire less yet are more sensitive to drought." Tree physiology 35, no. 5 (May 2015): 470-484. Full Text
  - Moshelion, M, Halperin, O, Wallach, R, Oren, R, and Way, DA. "Role of aquaporins in determining transpiration and photosynthesis in water-stressed plants: Crop water-use efficiency, growth and yield." Plant, Cell and Environment 38, no. 9 (2015): 1785-1793. Full Text
  - Tor-ngern, P, Oren, R, Ward, EJ, Palmroth, S, McCarthy, HR, and Domec, J-C. "Increases in atmospheric CO2 have little influence on transpiration of a temperate forest canopy." The New Phytologist 205, no. 2 (January 2015): 518-525. Full Text
  - De Kauwe, MG, Medlyn, BE, Zaehle, S, Walker, AP, Dietze, MC, Wang, Y-P, Luo, Y, Jain, AK, El-Masri, B, Hickler, T, Wårlind, D, Weng, E, Parton, WJ, Thornton, PE, Wang, S, Prentice, IC, Asao, S, Smith, B, McCarthy, HR, Iversen, CM, Hanson, PJ, Warren, JM, Oren, R, and Norby, RJ. "Where does the carbon go? A model-data intercomparison of vegetation carbon allocation and turnover processes at two temperate forest free-air CO2 enrichment sites." The New Phytologist 203, no. 3 (August 2014): 883-899. Full Text
  - Kim, D, Oren, R, Oishi, AC, Hsieh, C-I, Phillips, N, Novick, KA, and Stoy, PC. "Erratum to “Sensitivity of stand transpiration to wind velocity in a mixed broadleaved deciduous forest” [Agric. Forest Meteorol. 187 (2014) 62–71]." Agricultural and Forest Meteorology 192-193 (July 2014): 41-41. Full Text
  - Walker, AP, Hanson, PJ, De Kauwe, MG, Medlyn, BE, Zaehle, S, Asao, S, Dietze, M, Hickler, T, Huntingford, C, Iversen, CM, Jain, A, Lomas, M, Luo, Y, McCarthy, H, Parton, WJ, Prentice, IC, Thornton, PE, Wang, S, Wang, Y-P, Warlind, D, Weng, E, Warren, JM, Woodward, FI, Oren, R, and Norby, RJ. "Comprehensive ecosystem model-data synthesis using multiple data sets at two temperate forest free-air CO2enrichment experiments: Model performance at ambient CO2concentration." Journal of Geophysical Research: Biogeosciences 119, no. 5 (May 2014): 937-964. Full Text
  - Zaehle, S, Medlyn, BE, De Kauwe, MG, Walker, AP, Dietze, MC, Hickler, T, Luo, Y, Wang, Y-P, El-Masri, B, Thornton, P, Jain, A, Wang, S, Warlind, D, Weng, E, Parton, W, Iversen, CM, Gallet-Budynek, A, McCarthy, H, Finzi, A, Hanson, PJ, Prentice, IC, Oren, R, and Norby, RJ. "Evaluation of 11 terrestrial carbon-nitrogen cycle models against observations from two temperate Free-Air CO2 Enrichment studies." The New Phytologist 202, no. 3 (May 2014): 803-822. Full Text
  - Kim, D, Oren, R, Oishi, AC, Hsieh, C-I, Phillips, N, Novick, KA, and Stoy, PC. "Sensitivity of stand transpiration to wind velocity in a mixed broadleaved deciduous forest." Agricultural and Forest Meteorology 187 (April 2014): 62-71. Full Text
  - Oishi, AC, Palmroth, S, Johnsen, KH, McCarthy, HR, and Oren, R. "Sustained effects of atmospheric [CO2] and nitrogen availability on forest soil CO2 efflux." Glob Chang Biol 20, no. 4 (April 2014): 1146-1160. Full Text Link to Item
  - Oishi, AC, Palmroth, S, Johnsen, KH, Mccarthy, HR, and Oren, R. "Sustained effects of atmospheric [CO2] and nitrogen availability on forest soil CO2efflux." Global Change Biology 20, no. 4 (April 1, 2014): 1146-1160. Full Text
  - Nolan, RH, Mitchell, PJ, Bradstock, RA, and Lane, PNJ. "Structural adjustments in resprouting trees drive differences in post-fire transpiration." Tree physiology 34, no. 2 (February 16, 2014): 123-136. Full Text
  - De Kauwe, MG, Walker, AP, Dietze, MC, Hickler, T, Luo, Y, Wang, YP, El-Masri, B, Thornton, P, Jain, A, Wang, S, Warlind, D, Weng, E, Parton, W, Iversen, CM, Gallet-Budynek, A, Mccarthy, H, Finzi, A, Hanson, PJ, Prentice, IC, Oren, R, and Norby, RJ. "Evaluation of 11 terrestrial carbon-nitrogen cycle models against observations from two temperate Free-Air CO2 Enrichment studies." New Phytologist 202, no. 3 (January 1, 2014): 803-822. Full Text
  - Oishi, AC, Palmroth, S, Johnsen, KH, Mccarthy, HR, and Oren, R. "Sustained effects of atmospheric [CO2] and nitrogen availability on forest soil CO2 efflux." Global Change Biology 20, no. 4 (2014): 1146-1160.
  - De Kauwe, MG, Medlyn, BE, Zaehle, S, Walker, AP, Dietze, MC, Hickler, T, Jain, AK, Luo, Y, Parton, WJ, Prentice, IC, Smith, B, Thornton, PE, Wang, S, Wang, Y-P, Wårlind, D, Weng, E, Crous, KY, Ellsworth, DS, Hanson, PJ, Seok Kim, H, Warren, JM, Oren, R, and Norby, RJ. "Forest water use and water use efficiency at elevated CO2 : a model-data intercomparison at two contrasting temperate forest FACE sites." Global Change Biology 19, no. 6 (June 2013): 1759-1779. Full Text
  - Oishi, AC, Palmroth, S, Butnor, JR, Johnsen, KH, and Oren, R. "Spatial and temporal variability of soil CO2 efflux in three proximate temperate forest ecosystems." Agricultural and Forest Meteorology 171-172 (April 2013): 256-269. Full Text
  - Ward, EJ, Bell, DM, Clark, JS, and Oren, R. "Hydraulic time constants for transpiration of loblolly pine at a free-air carbon dioxide enrichment site." Tree Physiol 33, no. 2 (February 2013): 123-134. Full Text Link to Item
  - Ward, EJ, Oren, R, Bell, DM, Clark, JS, McCarthy, HR, Kim, H-S, and Domec, J-C. "The effects of elevated CO2 and nitrogen fertilization on stomatal conductance estimated from 11 years of scaled sap flux measurements at Duke FACE." Tree Physiology 33, no. 2 (February 2013): 135-151. Full Text
  - Kauwe, MGD, Medlyn, BE, Zaehle, S, Walker, AP, Dietze, MC, Hickler, T, Jain, AK, Luo, Y, Parton, WJ, Prentice, IC, Smith, B, Thornton, PE, Wang, S, Wang, Y-P, Wårlind, D, Weng, E, Crous, KY, Ellsworth, DS, Hanson, PJ, Kim, HS, Warren, JM, Oren, R, and Norby, RJ. "Forest water use and water use efficiency at elevated CO₂: A model-data intercomparison at two contrasting temperate forest FACE sites." Global Change Biology 19, no. 6 (2013): 1759-1779. Full Text
  - Drake, JE, Oishi, AC, Giasson, M-A, Oren, R, Johnsen, KH, and Finzi, AC. "Trenching reduces soil heterotrophic activity in a loblolly pine ( Pinus taeda ) forest exposed to elevated atmospheric [CO 2 ] and N fertilization." Agricultural and Forest Meteorology 165 (November 2012): 43-52. Full Text
  - Dieleman, WIJ, Vicca, S, Dijkstra, FA, Hagedorn, F, Hovenden, MJ, Larsen, KS, Morgan, JA, Volder, A, Beier, C, Dukes, JS, King, J, Leuzinger, S, Linder, S, Luo, Y, Oren, R, De Angelis, P, Tingey, D, Hoosbeek, MR, and Janssens, IA. "Simple additive effects are rare: a quantitative review of plant biomass and soil process responses to combined manipulations of CO2 and temperature." Global Change Biology 18, no. 9 (September 2012): 2681-2693. Full Text
  - Katul, GG, Oren, R, Manzoni, S, Higgins, C, and Parlange, MB. "Evapotranspiration: A process driving mass transport and energy exchange in the soil-plant-atmosphere-climate system." Reviews of Geophysics 50, no. 3 (September 2012). Full Text
  - Novick, KA, Katul, GG, McCarthy, HR, and Oren, R. "Increased resin flow in mature pine trees growing under elevated CO2 and moderate soil fertility." Tree Physiol 32, no. 6 (June 2012): 752-763. Full Text Link to Item
  - Bauerle, WL, Oren, R, Way, DA, Qian, SS, Stoy, PC, Thornton, PE, Bowden, JD, Hoffman, FM, and Reynolds, RF. "Photoperiodic regulation of the seasonal pattern of photosynthetic capacity and the implications for carbon cycling." Proceedings of the National Academy of Sciences of the United States of America 109, no. 22 (May 14, 2012): 8612-8617. Full Text
  - Long, X, Chen, C, Xu, Z, Oren, R, and He, J-Z. "Abundance and community structure of ammonia-oxidizing bacteria and archaea in a temperate forest ecosystem under ten-years elevated CO2." Soil Biology and Biochemistry 46 (March 2012): 163-171. Full Text
  - Zhang, Y, Oren, R, and Kang, S. "Spatiotemporal variation of crown-scale stomatal conductance in an arid Vitis vinifera L. cv. Merlot vineyard: direct effects of hydraulic properties and indirect effects of canopy leaf area." Tree Physiology 32, no. 3 (March 2012): 262-279. Full Text
  - Ellsworth, DS, Thomas, R, Crous, KY, Palmroth, S, Ward, E, Maier, C, DeLucia, E, and Oren, R. "Elevated CO2 affects photosynthetic responses in canopy pine and subcanopy deciduous trees over 10 years: a synthesis from Duke FACE." Global Change Biology 18, no. 1 (January 2012): 223-242. Full Text
  - Oren, R, and Bird, C. "Message from the editor-in-chief." Tree Physiology 32, no. 2 (2012): i-. Full Text
  - Lee, X, Goulden, ML, Hollinger, DY, Barr, A, Black, TA, Bohrer, G, Bracho, R, Drake, B, Goldstein, A, Gu, L, Katul, G, Kolb, T, Law, BE, Margolis, H, Meyers, T, Monson, R, Munger, W, Oren, R, Paw U, KT, Richardson, AD, Schmid, HP, Staebler, R, Wofsy, S, and Zhao, L. "Observed increase in local cooling effect of deforestation at higher latitudes." Nature 479, no. 7373 (November 16, 2011): 384-387. Full Text
  - Bertin, S, Palmroth, S, Kim, HS, Perks, MP, Mencuccini, M, and Oren, R. "Modelling understorey light for seedling regeneration in continuous cover forestry canopies." Forestry 84, no. 4 (October 1, 2011): 397-409. Full Text
  - Weng, E, Luo, Y, Gao, C, and Oren, R. "Uncertainty analysis of forest carbon sink forecast with varying measurement errors: a data assimilation approach." Journal of Plant Ecology 4, no. 3 (September 2011): 178-191. Full Text
  - Drake, JE, Gallet-Budynek, A, Hofmockel, KS, Bernhardt, ES, Billings, SA, Jackson, RB, Johnsen, KS, Lichter, J, McCarthy, HR, McCormack, ML, Moore, DJP, Oren, R, Palmroth, S, Phillips, RP, Pippen, JS, Pritchard, SG, Treseder, KK, Schlesinger, WH, Delucia, EH, and Finzi, AC. "Increases in the flux of carbon belowground stimulate nitrogen uptake and sustain the long-term enhancement of forest productivity under elevated CO₂." Ecology Letters 14, no. 4 (April 2011): 349-357. Full Text
  - Hollinger, DY, Wharton, S, Oren, R, Noormets, A, Fischer, ML, Verma, SB, Cook, DR, Sun, G, McNulty, S, Wofsy, SC, Bolstad, PV, Burns, SP, Curtis, PS, Drake, BG, Falk, M, Foster, DR, Gu, L, Hadley, JL, Katul, GG, Litvak, M, Ma, S, Martin, TA, Matamala, R, Meyers, TP, Monson, RK, Munger, JW, Oechel, WC, Paw, UKT, Schmid, HP, Scott, RL, Starr, G, Suyker, AE, and Torn, MS. "Assessing net ecosystem carbon exchange of U.S. terrestrial ecosystems by integrating eddy covariance flux measurements and satellite observations." Agricultural and Forest Meteorology 151, no. 1 (January 15, 2011): 60-69. Full Text
  - Kim, H-S, Palmroth, S, Thérézien, M, Stenberg, P, and Oren, R. "Analysis of the sensitivity of absorbed light and incident light profile to various canopy architecture and stand conditions." Tree Physiol 31, no. 1 (January 2011): 30-47. Full Text Link to Item
  - Oren, R, and Kennedy, C. "Introduction." Tree Physiology 31, no. 1 (January 1, 2011): i-i. Full Text
  - Oren, R, and Kennedy, C. "Changes implemented in Tree Physiology." Tree physiology 31, no. 1 (2011): i-. Full Text
  - Warren, JM, Norby, RJ, Wullschleger, SD, and Oren, R. "Elevated CO₂ enhances leaf senescence during extreme drought in a temperate forest." Tree Physiology 31, no. 2 (2011): 117-130. Full Text
  - Ge, Y, Chen, C, Xu, Z, Oren, R, and He, J-Z. "The spatial factor, rather than elevated CO₂, controls the soil bacterial community in a temperate Forest Ecosystem." Applied and Environmental Microbiology 76, no. 22 (November 2010): 7429-7436. Full Text
  - Palmroth, S, Katul, GG, Hui, D, McCarthy, HR, Jackson, RB, and Oren, R. "Estimation of long-term basin scale evapotranspiration from streamflow time series." Water Resources Research 46, no. 10 (October 2010). Full Text Open Access Copy
  - Domec, J-C, Schäfer, K, Oren, R, Kim, HS, and McCarthy, HR. "Variable conductivity and embolism in roots and branches of four contrasting tree species and their impacts on whole-plant hydraulic performance under future atmospheric CO₂ concentration." Tree Physiology 30, no. 8 (August 2010): 1001-1015. Full Text
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  - Oren, R, and Pataki, DE. "Transpiration in response to variation in microclimate and soil moisture in southeastern deciduous forests." Oecologia 127, no. 4 (2001): 549-559. Full Text
  - Phillips, N, Bergh, J, Oren, R, and Linder, S. "Effects of nutrition and soil water availability on water use in a Norway spruce stand." Tree Physiology 21, no. 12-13 (2001): 851-860.
  - Schulze, W, Schulze, ED, Schulze, I, and Oren, R. "Quantification of insect nitrogen utilization by the venus fly trap Dionaea muscipula catching prey with highly variable isotope signatures." Journal of Experimental Botany 52, no. 358 (2001): 1041-1049.
  - "Modeling CO2 and water vapor turbulent flux distributions within a forest canopy." Journal of Geophysical Research Atmospheres 105, no. D21 (November 16, 2000): 26333-26351. Full Text
  - Ewers, BE, Oren, R, and Sperry, JS. "Influence of nutrient versus water supply on hydraulic architecture and water balance in Pinus taeda." Plant, Cell and Environment 23, no. 10 (October 2000): 1055-1066. Full Text
  - Hacke, UG, Sperry, JS, Ewers, BE, Ellsworth, DS, Schäfer, KVR, and Oren, R. "Influence of soil porosity on water use in Pinus taeda." Oecologia 124, no. 4 (September 2000): 495-505. Full Text
  - Ewers, BE, and Oren, R. "Analyses of assumptions and errors in the calculation of stomatal conductance from sap flux measurements." Tree Physiol 20, no. 9 (May 2000): 579-589. Link to Item
  - Lai, C-T, Katul, G, Ellsworth, D, and Oren, R. "Modelling Vegetation-Atmosphere Co2 Exchange By A Coupled Eulerian-Langrangian Approach." Boundary Layer Meteorology 95, no. 1 (April 2000): 91-122. Full Text
  - Schafer, KVR, Oren, R, and Tenhunen, JD. "The effect of tree height on crown level stomatal conductance." Plant, Cell and Environment 23, no. 4 (April 2000): 365-375. Full Text
  - "Sap flux of co-occurring species in a western subalpine forest during seasonal soil drought." Ecology 81, no. 9 (January 1, 2000): 2557-2566. Full Text
  - "Survey and synthesis of intra- and interspecific variation in stomatal sensitivity to vapour pressure deficit." Plant, Cell and Environment 22, no. 12 (December 1, 1999): 1515-1526. Full Text
  - "Temporal patterns of water flux in trees and lianas in a Panamanian moist forest." Trees Structure and Function 14, no. 3 (November 1, 1999): 116-123.
  - Katul, G, Hsieh, C-I, Bowling, D, Clark, K, Shurpali, N, Turnipseed, A, Albertson, J, Tu, K, Hollinger, D, Evans, B, Offerle, B, Anderson, D, Ellsworth, D, Vogel, C, and Oren, R. "Spatial Variability of Turbulent Fluxes in the Roughness Sublayer of an Even-Aged Pine Forest." Boundary Layer Meteorology 93, no. 1 (October 1999): 1-28. Full Text
  - Oren, R, Phillips, N, Ewers, BE, Pataki, DE, and Megonigal, JP. "Sap-flux-scaled transpiration responses to light, vapor pressure deficit, and leaf area reduction in a flooded Taxodium distichum forest." Tree Physiology 19, no. 6 (May 1999): 337-347. Full Text
  - "Carry-over effects of water and nutrient supply on water use of Pinus taeda." Ecological Applications 9, no. 2 (January 1, 1999): 513-525. Full Text
  - Pataki, DE, Oren, R, Katul, G, and Sigmon, J. "Canopy conductance of Pinus taeda, Liquidambar styraciflua and Quercus phellos under varying atmospheric and soil water conditions." Tree Physiology 18, no. 5 (May 1998): 307-315. Full Text
  - Pataki, DE, Oren, R, and Phillips, N. "Responses of sap flux and stomatal conductance of Pinus taeda L. trees to stepwise reductions in leaf area." Journal of Experimental Botany 49, no. 322 (May 1, 1998): 871-878. Full Text
  - "Water balance delineates the soil layer in which moisture affects canopy conductance." Ecological Applications 8, no. 4 (January 1, 1998): 990-1002. Full Text
  - Oren, R, Phillips, N, Katul, G, Ewers, BE, and Pataki, DE. "Scaling xylem sap flux and soil water balance and calculating variance: a method for partitioning water flux in forests." Annales Des Sciences Forestières 55, no. 1-2 (1998): 191-216. Full Text
  - Pataki, DE, Oren, R, and Tissue, DT. "Elevated carbon dioxide does not affect average canopy stomatal conductance of Pinus taeda L." Oecologia 117, no. 1-2 (1998): 47-52. Full Text
  - Phillips, N, and Oren, R. "A comparison of daily representations of canopy conductance based on two conditional time-averaging methods and the dependence of daily conductance on environmental factors." Annales Des Sciences Forestières 55, no. 1-2 (1998): 217-235. Full Text
  - "Winter and spring thaw as observed with imaging radar at BOREAS." Journal of Geophysical Research Atmospheres 102, no. 24 (December 26, 1997): 29673-29684.
  - Hogg, EH, Black, TA, den Hartog, G, Neumann, HH, Zimmermann, R, Hurdle, PA, Blanken, PD, Nesic, Z, Yang, PC, Staebler, RM, McDonald, KC, and Oren, R. "A comparison of sap flow and eddy fluxes of water vapor from a boreal deciduous forest." Journal of Geophysical Research: Atmospheres 102, no. D24 (December 26, 1997): 28929-28937. Full Text
  - Way, J, Zimmermann, R, Rignot, E, McDonald, K, and Oren, R. "Winter and spring thaw as observed with imaging radar at BOREAS." Journal of Geophysical Research: Atmospheres 102, no. D24 (December 26, 1997): 29673-29684. Full Text
  - Zimmermann, R, Hurdle, PA, Blanken, PD, Nesic, Z, Yang, PC, Staebler, RM, McDonald, KC, and Oren, R. "A comparison of sap flow and eddy fluxes of water vapor from a boreal deciduous forest." Journal of Geophysical Research Atmospheres 102, no. 24 (December 26, 1997): 28929-28937. (Review)
  - Hsieh, CI, Phillips, N, and Lewin, K. "Lagrangian dispersion model for predicting CO<inf>2</inf> sources, sinks, and fluxes in a uniform loblolly pine (Pinus taeda L.) stand." Journal of Geophysical Research Atmospheres 102, no. 8 (April 27, 1997): 9309-9321.
  - Katul, G, Oren, R, Ellsworth, D, Hsieh, C-I, Phillips, N, and Lewin, K. "A Lagrangian dispersion model for predicting CO2sources, sinks, and fluxes in a uniform loblolly pine (Pinus taeda L.) stand." Journal of Geophysical Research: Atmospheres 102, no. D8 (April 27, 1997): 9309-9321. Full Text
  - Katul, G, Todd, P, Pataki, D, Kabala, ZJ, and Oren, R. "Soil water depletion by oak trees and the influence of root water uptake on the moisture content spatial statistics." Water Resources Research 33, no. 4 (April 1997): 611-623. Full Text
  - Phillips, N, Nagchaudhuri, A, Oren, R, and Katul, G. "Time constant for water transport in loblolly pine trees estimated from time series of evaporative demand and stem sapflow." Trees 11, no. 7 (1997): 412-412. Full Text
  - "Latent and sensible heat flux predictions from a uniform pine forest using surface renewal and flux variance methods." Boundary Layer Meteorology 80, no. 1-2 (December 1, 1996): 249-282.
  - Katul, G, Hsieh, CI, Oren, R, Ellsworth, D, and Phillips, N. "Latent and sensible heat flux predictions from a uniform pine forest using surface renewal and flux variance methods." BOUNDARY-LAYER METEOROLOGY 80, no. 3 (August 1996): 249-282. Full Text Link to Item
  - Ellsworth, DS, Oren, R, Huang, C, Phillips, N, and Hendrey, GR. "Leaf and canopy responses to elevated CO2 in a pine forest under free-air CO2 enrichment (vol 104, pg 139, 1995)." OECOLOGIA 106, no. 3 (May 1996): 416-416. Full Text Link to Item
  - Oren, R, Zimmermann, R, and Terbough, J. "Transpiration in Upper Amazonia Floodplain and Upland Forests in Response to Drought-Breaking Rains." Ecology 77, no. 3 (April 1996): 968-973. Full Text
  - "Radial patterns of xylem sap flow in non-, diffuse- and ring-porous tree species." Plant, Cell and Environment 19, no. 8 (January 1, 1996): 983-990. Full Text
  - "Erratum: Leaf and canopy responses to elevated CO2 in a pine forest under free-air CO2 enrichment (Oecologia (1995) 104 (139-146))." Oecologia 106, no. 3 (January 1, 1996): 416-null.
  - WARING, RH, WAY, JB, HUNT, ER, MORRISSEY, L, RANSON, KJ, WEISHAMPEL, JF, OREN, R, and FRANKLIN, SE. "BIOLOGISTS TOOLBOX - IMAGING RADAR FOR ECOSYSTEM STUDIES." Bioscience 45, no. 10 (November 1, 1995): 715-723. (Academic Article) Full Text Link to Item
  - Waring, RH, Way, J, Hunt,, ER, Morrissey, L, Ranson, KJ, Weishampel, JF, Oren, R, and Franklin, SE. "Imaging Radar for Ecosystem Studies." Bioscience 45, no. 10 (November 1995): 715-723. Full Text
  - "Dielectric and hydraulic response of selected forest canopies at the boreas test sites in Canada." International Geoscience and Remote Sensing Symposium (Igarss) 2 (January 1, 1995): 1204-1206.
  - Buchmann, N, Oren, R, and Zimmermann, R. "Response of magnesium-deficient saplings in a young, open stand of Picea abies (L.) Karst. to elevated soil magnesium, nitrogen and carbon." Environmental Pollution 87, no. 1 (1995): 31-43. Full Text
  - Ellsworth, DS, Oren, R, Huang, C, Phillips, N, and Hendrey, GR. "Leaf and canopy responses to elevated CO₂ in a pine forest under free-air CO₂ enrichment." Oecologia 104, no. 2 (1995): 139-146. Full Text
  - Zimmermann, R, McDonald, K, Oren, R, and Way, J. "Xylem dielectric constant, water status, and transpiration of young Jack Pine (Pinus banksiana Lamb.) in the southern boreal zone of Canada." International Geoscience and Remote Sensing Symposium (IGARSS) 2 (1995): 1006-1008.
  - "Microclimate, water potential, transpiration, and bole dielectric constant of coniferous and deciduous tree species in the continental boreal ecotone of central Alaska." International Geoscience and Remote Sensing Symposium (Igarss) 1 (December 1, 1994): 226-228.
  - Way, J, Rignot, EJM, McDonald, KC, Oren, R, Kwok, R, Bonan, G, Dobson, MC, Viereck, LA, and Roth, JE. "Evaluating the type and state of Alaska taiga forests with imaging radar for use in ecosystem models." Ieee Transactions on Geoscience and Remote Sensing 32, no. 2 (March 1994): 353-370. Full Text
  - Berger, A, Oren, R, and Schulze, E-D. "Element concentrations in the xylem sap of Picea abies (L.) Karst. seedlings extracted by various methods under different environmental conditions." Tree Physiology 14, no. 2 (February 1, 1994): 111-128. Full Text
  - "Hydrologic and dielectric properties of woody plant tissue: Implications for remote sensing of canopy water status." International Geoscience and Remote Sensing Symposium (Igarss) 4 (December 1, 1993): 2062-null.
  - Oren, R, Werk, KS, Buchmann, N, and Zimmermann, R. "Chlorophyll–nutrient relationships identify nutritionally caused decline in Piceaabies stands." Canadian Journal of Forest Research 23, no. 6 (June 1993): 1187-1195. Full Text
  - BUCHMANN, N, OREN, R, GEBAUER, G, DIETRICH, P, and SCHULZE, ED. "THE USE OF STABLE ISOTOPES IN ECOSYSTEM RESEARCH - 1ST RESULTS OF A FIELD-STUDY WITH N-15." ISOTOPENPRAXIS 28, no. 1 (1992): 51-59. Full Text Link to Item
  - Katz, C, Oren, R, Schulze, E-D, and Milburn, JA. "Uptake of water and solutes through twigs of Picea abies (L.) Karst." Trees 3, no. 1 (May 1989). Full Text
  - Oren, R, Werk, KS, Schulze, E-D, Meyer, J, Schneider, BU, and Schramel, P. "Performance of two Picea abies (L.) Karst. stands at different stages of decline." Oecologia 78, no. 4 (1989): 571-571. Full Text
  - Oren, R, Schulze, E-D, Werk, KS, and Meyer, J. "Performance of two Picea abies (L.) Karst. stands at different stages of decline : VII. Nutrient relations and growth." Oecologia 77, no. 2 (November 1988): 163-173. Full Text
  - Meyer, J, Schneider, BU, Werk, K, Oren, R, and Schulze, E-D. "Performance of two Picea abies (L.) Karst. stands at different stages of decline." Oecologia 77, no. 1 (October 1988): 7-13. Full Text
  - Osonubi, O, Oren, R, Werk, KS, Schulze, E-D, and Heilmeier, H. "Performance of two Picea abies (L.) Karst. stands at different stages of decline." Oecologia 77, no. 1 (October 1988): 1-6. Full Text
  - Werk, KS, Oren, R, Schulze, E-D, Zimmermann, R, and Meyer, J. "Performance of two Picea abies (L.) Karst. stands at different stages of decline." Oecologia 76, no. 4 (September 1988): 519-524. Full Text
  - Zimmermann, R, Oren, R, Schulze, E-D, and Werk, KS. "Performance of two Picea abies (L.) Karst. stands at different stages of decline." Oecologia 76, no. 4 (September 1988): 513-518. Full Text
  - Oren, R, Schulze, E-D, Werk, KS, Meyer, J, Schneider, BU, and Heilmeier, H. "Performance of two Picea abies (L.) Karst. stands at different stages of decline - I. Carbon relations and stand growth." Oecologia 75, no. 1 (1988): 25-37. Full Text
  - Oren, R, Werk, KS, Schulze, E-D, Meyer, J, Schneider, BU, and Schramel, P. "Performance of two Picea abies (L.) Karst. stands at different stages of decline - VI. Nutrient concentration." Oecologia 77, no. 2 (1988): 151-162. Full Text
  - STEUDLE, E, OREN, R, and SCHULZE, ED. "WATER TRANSPORT IN MAIZE ROOTS - MEASUREMENT OF HYDRAULIC CONDUCTIVITY, SOLUTE PERMEABILITY, AND OF REFLECTION COEFFICIENTS OF EXCISED ROOTS USING THE ROOT PRESSURE PROBE." PLANT PHYSIOLOGY 84, no. 4 (August 1987): 1220-1232. (Academic Article) Full Text
  - OREN, R, WARING, RH, STAFFORD, SG, and BARRETT, JW. "24 YEARS OF PONDEROSA PINE GROWTH IN RELATION TO CANOPY LEAF-AREA AND UNDERSTORY COMPETITION." FOREST SCIENCE 33, no. 2 (June 1987): 538-547. Link to Item
  - "Twenty-four years of ponderosa pine growth in relation to canopy leaf area and understory competition." Forest Science 33, no. 2 (January 1, 1987): 538-547.
  - Oren, R, Schulze, E-D, Matyssek, R, and Zimmermann, R. "Estimating photosynthetic rate and annual carbon gain in conifers from specific leaf weight and leaf biomass." Oecologia 70, no. 2 (September 1986): 187-193. Full Text
  - Oren, R, Werk, KS, and Schulze, E-D. "Relationships between foliage and conducting xylem in Picea abies (L.) Karst." Trees 1, no. 1 (1986). Full Text
  - Oren, R, Thies, WG, and Waring, RH. "Tree vigor and stand growth of Douglas-fir as influenced by laminated root rot." Canadian Journal of Forest Research 15, no. 5 (October 1985): 985-988. Full Text
  - Waring, RH, and Barrett, JW. "Attacks of mountain pine beetle as related to tree vigor of ponderosa pine." Forest Science 29, no. 2 (January 1, 1983): 395-402.
  - Waring, RH, Schroeder, PE, and Oren, R. "Application of the pipe model theory to predict canopy leaf area." Canadian Journal of Forest Research 12, no. 3 (September 1982): 556-560. Full Text
  - Song, C, Katul, G, Oren, R, Band, LE, Tague, CL, Stoy, PC, and McCarthy, HR. "Energy, water, and carbon fluxes in a loblolly pine stand: Results from uniform and gappy canopy models with comparisons to eddy flux data(Published online)." Journal of Geophysical Research 114, no. G4. Full Text Open Access Copy
  - Way, DA, Oren, R, Kim, H-S, and Katul, GG. "How well do stomatal conductance models perform on closing plant carbon budgets? A test using seedlings grown under current and elevated air temperatures(Published online)." Journal of Geophysical Research 116, no. G4. Full Text
- Book Sections
  - Luxmoore, RJ, Oren, R, Sheriff, DW, and Thomas, RB. "Source–Sink–Storage Relationships of Conifers." In Resource Physiology of Conifers, 179-216.: Elsevier, 1995. Full Text
  - Margolis, H, Oren, R, Whitehead, D, and Kaufmann, MR. "Leaf Area Dynamics of Conifer Forests." In Ecophysiology of Coniferous Forests, 181-223.: Elsevier, 1995. Full Text
  - Matyssek, R, Reich, P, Oren, R, and Winner, WE. "Response Mechanisms of Conifers to Air Pollutants." In Ecophysiology of Coniferous Forests, 255-308.: Elsevier, 1995. Full Text
  - Oren, R, and Sheriff, DW. "Water and Nutrient Acquisition by Roots and Canopies." In Resource Physiology of Conifers, 39-74.: Elsevier, 1995. Full Text
- Conference Papers
  - McDonald, KC, Zimmerman, R, and Way, J. "An Investigation Of The Relationship Between Tree Water Potential And Dielectric Constant(Accepted)." Geoscience and Remote Sensing Symposium, 1992. IGARSS '92. International.: IEEE,. Full Text
Teaching & Mentoring
Recent Courses
- ENVIRON 393: Research Independent Study 2019
- ENVIRON 503: Forest Ecosystems 2019
- ENVIRON 593: Independent Studies and Projects 2019
- ENVIRON 898: Program Area Seminar 2019
- ENVIRON 899: Master's Project 2019

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