Education
University of Pennsylvania ·
1979
Ph.D.
University of Pennsylvania ·
1973
B.A.
Training & Certifications
Student ·
2009 - 2009
University of Wisconsin, Madison
, Summer institute for scientific teaching
Associate Professor ·
1988 - 1992
Harvard University
, Department of cellular and developmental biology
Student ·
1985 - 1985
Marine Biological Laboratory
, Physiology course
Assistant Professor ·
1984 - 1988
Harvard University
, Department of cellular and developmental biology
Research Associate (Research Faculty) ·
1982 - 1984
Johns Hopkins University, School of Medicine
Postdoctoral Fellow ·
1978 - 1982
Johns Hopkins University
, Cell biology and anatomy