Kathleen M. Pryer | Scholars@Duke

Kathleen M. Pryer
Professor in the Department of Biology

My research focuses on understanding the evolutionary relationships of ancient land plants, especially ferns and horsetails, by integrating evidence from morphology, molecules (DNA sequence data from multiple genes), and the fossil record. I use an explicit phylogenetic framework to examine the morphological evolution of various sporophytic and gametophytic characters within vascular plants, and to gain insight into the evolution of various life history traits and the body plans that typify vascular plants. A phylogenetic perspective also informs my molecular evolutionary studies that attempt to elucidate why we observe remarkable rate heterogeneity in chloroplast genes in land plant phylogeny.

Current Appointments & Affiliations

Professor in the Department of Biology, Biology, Trinity College of Arts & Sciences 2011
Associate of the Duke Initiative for Science & Society, Duke Science & Society, Initiatives 2017

Contact Information

358 Bio Sci Bldg, Durham, NC 27708
Box 90338, Durham, NC 27708-0338
(919) 660-7380
Pryer Lab Website

Background
Education, Training, & Certifications
- Ph.D., Duke University 1995
- M.S., University of Guelph (Canada) 1981
- B.S., McGill University (Canada) 1976
Duke Appointment History
- Associate Professor of Biology, Biology, Trinity College of Arts & Sciences 2005 - 2011
- Assistant Professor of Biology, Biology, Trinity College of Arts & Sciences 2001 - 2004
Recognition
In the News
- JAN 9, 2018 Duke Research Blog
  
  Kathleen Pryer: A Passion for the Little-Loved Fern
- JUL 2, 2015 Nature
  
  Kathleen Pryer comments: Plant collections left in the cold by cuts
- FEB 16, 2015 redOrbit
  
  Distant fern species create 'love child' after 60 million years
- FEB 13, 2015
  
  Distant Species Produce Love Child After 60-Millon-Year Breakup
- JUL 18, 2014 Durham Herald-Sun
  
  Duke researcher uses crowdfunding to raise money for fern genome project
- JUL 14, 2014 Scientific American
  
  How scientists uncovered Arctic clues to a past where a tiny fern changed the planet
- JUL 2, 2014
  
  Crowdfunding for Azolla Fern Research Hits Target
- JUN 20, 2014 The Economist
  
  Finding the genome of an extraordinary plant
- JUN 19, 2014
  
  Going Public With Research Funding
- JUN 13, 2014 Phys.org
  
  Tiny plant, big potential: Researchers raise money from public to sequence 'superorganism' genome
- MAY 30, 2014 The Globe and Mail
  
  Kathleen Pryer: This lowly fern has massive green potential
Awards & Honors
- Faculty Early Career Development (CAREER) Program. National Science Foundation. 2004
Expertise
Global Scholarship
- Research
  - Costa Rica (Country)
  - Mexico (Country)
Research
Selected Grants
- Oceans Apart: Assessing the roles of Vicariance and Long-distance Dispersal in the Widely Disjunct Fern Genus Cheilanthes (Pteridaceae) awarded by National Geographic Society 2019
- DISSERTATION RESEARCH: Evolution and Function of Farina in the Desiccation-tolerant Notholaenid Ferns (Pteridaceae) awarded by National Science Foundation 2017 - 2019
- Collaborative Research: Digitization TCN: The Macroalgal Herbarium Consortium: awarded by University of North Carolina - Chapel Hill 2013 - 2018
- Identifying those Pesky Mustards: The Boechera Digitization Project awarded by Andrew W. Mellon Foundation 2014 - 2017
- DISSERTATION RESEARCH: Early Evolution of Fungi and the Transition to a Terrestrial Lifestyle: Marine Habitats, the New awarded by National Science Foundation 2013 - 2017
- Collaborative Research: Exploring the Evolution of an Extreme Makeover in Ferns awarded by National Science Foundation 2012 - 2016
- DISSERTATION RESEARCH: Assessing the Evolutionary Significance of Recurrent Horizontal Gene Transfer of a Chimeric Phot awarded by National Science Foundation 2014 - 2016
- DISSERTATION RESEARCH: Untangling the Cystopteris fragilis Complex: An Investigation of the Effects of Polploidy on Div awarded by National Science Foundation 2011 - 2015
- DISSERTATION RESEARCH: Patterns of Gene Expression in Allopolyploid Taxa of Recurrent Origin: A Case Study of the Polypo awarded by National Science Foundation 2011 - 2014
- DISSERTATION RESEARCH: Subsexual Processes and the Origin of Genetic Variation in Asexual Ferns awarded by National Science Foundation 2011 - 2014
- Collaborative Research: The Roles of Polyploidy and Apomixis in Shaping the Diversity and Distribution of Cheilanthoid F awarded by National Science Foundation 2007 - 2011
- A plan to rehouse and reorganize the vascular plant herbarium and botanical library at Duke University (DUKE) awarded by National Science Foundation 2007 - 2011
- Career: Reconciling patterns of phylogenetic rate heterogenity in ferns with morphology, ecology and life history awarded by National Science Foundation 2004 - 2010
- DISSERTATION RESEARCH: Evolution and diversification of epiphytic ferns awarded by National Science Foundation 2004 - 2007
- REU Site: Bioinformatic and Phylogenetic approaches to the study of plant and fungal biodiversity awarded by National Science Foundation 2004 - 2007
- Collaborative Research: Using phylogeny to investigate the history of morphological evolution in heterosporous ferns awarded by National Science Foundation 2001 - 2005
- (93-0602) Ontogeny and Phylogeny in the Aquatic Fern Family Marsileaceae awarded by National Science Foundation 1993 - 1995
Publications & Artistic Works
Selected Publications
- Book Sections
  - Schuettpelz, E, and Pryer, KM. "Fern phylogeny." Biology and Evolution of Ferns and Lycophytes. January 1, 2008. 395-416. Full Text
  - Smith, AR, Pryer, KM, Schuettpelz, E, Korall, P, Schneider, H, and Wolf, PG. "Fern classification." Biology and Evolution of Ferns and Lycophytes. January 1, 2008. 417-467. Full Text
- Conference Papers
  - Windham, MD, Huiet, L, Schuettpelz, E, Grusz, AL, Rothfels, C, Beck, J, Yatskievych, G, and Pryer, KM. "Using Plastid and Nuclear DNA Sequences to Redraw Generic Boundaries and Demystif Species Complexes in Cheilanthoid Ferns." 2009. Link to Item
  - Pryer, KM, Schneider, H, and Magallon, S. "The radiation of vascular plants." 2004. Link to Item
- Journal Articles
  - Dijkhuizen, LW, Brouwer, P, Bolhuis, H, Reichart, G-J, Koppers, N, Huettel, B, Bolger, AM, Li, F-W, Cheng, S, Liu, X, Wong, GK-S, Pryer, K, Weber, A, Bräutigam, A, and Schluepmann, H. "Is there foul play in the leaf pocket? The metagenome of floating fern Azolla reveals endophytes that do not fix N2 but may denitrify." The New phytologist 217.1 (January 2018): 453-466. Full Text
  - Song, M, Kuo, L-Y, Huiet, L, Pryer, KM, Rothfels, CJ, and Li, F-W. "A novel chloroplast gene reported for flagellate plants." American journal of botany 105.1 (January 2018): 117-121. Full Text
  - Rothfels, CJ, Pryer, KM, and Li, F-W. "Next-generation polyploid phylogenetics: rapid resolution of hybrid polyploid complexes using PacBio single-molecule sequencing." The New phytologist 213.1 (January 2017): 413-429. Full Text
  - Hirai, R, Schuettpelz, E, Huiet, L, Pryer, K, Smith, A, and Prado, J. "Phylogeny and relationships of the neotropical Adiantum raddianum group (Pteridaceae)." Taxon 65.6 (December 1, 2016): 1225-1235. Full Text
  - Haufler, CH, Pryer, KM, Schuettpelz, E, Sessa, EB, Farrar, DR, Moran, R, Schneller, JJ, Watkins, JE, and Windham, MD. "Sex and the Single Gametophyte: Revising the Homosporous Vascular Plant Life Cycle in Light of Contemporary Research." BioScience 66.11 (November 1, 2016): 928-937. Full Text Open Access Copy
  - Schuettpelz, E, Chen, C-W, Kessler, M, Pinson, JB, Johnson, G, Davila, A, Cochran, AT, Huiet, L, and Pryer, KM. "A revised generic classification of vittarioid ferns (Pteridaceae) based on molecular, micromorphological, and geographic data." Taxon 65.4 (August 30, 2016): 708-722. Full Text
  - Li, F-W, Kuo, L-Y, Pryer, KM, and Rothfels, CJ. "Genes Translocated into the Plastid Inverted Repeat Show Decelerated Substitution Rates and Elevated GC Content." Genome Biology and Evolution 8.8 (August 2016): 2452-2458. Full Text
  - Pryer, KM, Huiet, L, Li, F-W, Rothfels, CJ, and Schuettpelz, E. "Maidenhair Ferns, Adiantum , are Indeed Monophyletic and Sister to Shoestring Ferns, Vittarioids (Pteridaceae)." Systematic Botany 41.1 (March 1, 2016): 17-23. Full Text
  - Grusz, AL, and Pryer, KM. "Development of microsatellite markers for the apomictic triploid fern Myriopteris lindheimeri (Pteridaceae)." Applications in plant sciences 3.11 (November 10, 2015). Full Text
  - Kao, T-T, Pryer, KM, Turner, MD, White, RA, and Korall, P. "Origins of the Endemic Scaly Tree Ferns on the Galápagos and Cocos Islands." International Journal of Plant Sciences 176.9 (November 2015): 869-879. Full Text
  - Schuettpelz, E, Pryer, KM, and Windham, MD. "A Unified Approach to Taxonomic Delimitation in the Fern Genus Pentagramma (Pteridaceae)." Systematic Botany 40.3 (October 1, 2015): 629-644. Full Text Open Access Copy
  - Wolf, PG, Sessa, EB, Marchant, DB, Li, F-W, Rothfels, CJ, Sigel, EM, Gitzendanner, MA, Visger, CJ, Banks, JA, Soltis, DE, Soltis, PS, Pryer, KM, and Der, JP. "An Exploration into Fern Genome Space." Genome Biology and Evolution 7.9 (September 2015): 2533-2544. Full Text Open Access Copy
  - Rothfels, CJ, Li, F-W, Sigel, EM, Huiet, L, Larsson, A, Burge, DO, Ruhsam, M, Deyholos, M, Soltis, DE, Stewart, CN, Shaw, SW, Pokorny, L, Chen, T, dePamphilis, C, DeGironimo, L, Chen, L, Wei, X, Sun, X, Korall, P, Stevenson, DW, Graham, SW, Wong, GK-S, and Pryer, KM. "The evolutionary history of ferns inferred from 25 low-copy nuclear genes." American journal of botany 102.7 (July 16, 2015): 1089-1107. Full Text Open Access Copy
  - Rothfels, CJ, Johnson, AK, Hovenkamp, PH, Swofford, DL, Roskam, HC, Fraser-Jenkins, CR, Windham, MD, and Pryer, KM. "Natural hybridization between genera that diverged from each other approximately 60 million years ago." The American naturalist 185.3 (March 2015): 433-442. Full Text Open Access Copy
  - Huiet, L, Lenz, M, Nelson, JK, Pryer, KM, and Smith, AR. "Adiantumshastense, a new species of maidenhair fern from California." PhytoKeys 53 (January 2015): 73-81. Full Text Open Access Copy
  - Li, F-W, Melkonian, M, Rothfels, CJ, Villarreal, JC, Stevenson, DW, Graham, SW, Wong, GK-S, Pryer, KM, and Mathews, S. "Phytochrome diversity in green plants and the origin of canonical plant phytochromes." Nature communications 6 (January 2015): 7852-. Full Text Open Access Copy
  - Li, F-W, Rothfels, CJ, Melkonian, M, Villarreal, JC, Stevenson, DW, Graham, SW, Wong, GK-S, Mathews, S, and Pryer, KM. "The origin and evolution of phototropins." Frontiers in plant science 6 (January 2015): 637-. Full Text Open Access Copy
  - Rothfels, CJ, Johnson, AK, Windham, MD, and Pryer, KM. "Low-copy nuclear data confirm rampant allopolyploidy in the Cystopteridaceae (Polypodiales)." Taxon 63.5 (October 29, 2014): 1026-1036. Full Text
  - Sigel, EM, Windham, MD, Haufler, CH, and Pryer, KM. "Phylogeny, Divergence Time Estimates, and Phylogeography of the Diploid Species of the Polypodium vulgare Complex (Polypodiaceae)." Systematic Botany 39.4 (October 1, 2014): 1042-1055. Full Text
  - Sigel, EM, Windham, MD, and Pryer, KM. "Evidence for reciprocal origins in Polypodium hesperium (Polypodiaceae): a fern model system for investigating how multiple origins shape allopolyploid genomes." American journal of botany 101.9 (September 17, 2014): 1476-1485. Full Text
  - Grusz, AL, Windham, MD, Yatskievych, G, Huiet, L, Gastony, GJ, and Pryer, KM. "Patterns of Diversification in the Xeric-adapted Fern Genus Myriopteris (Pteridaceae)." Systematic Botany 39.3 (July 1, 2014): 698-714. Full Text
  - Li, F-W, Villarreal, JC, Kelly, S, Rothfels, CJ, Melkonian, M, Frangedakis, E, Ruhsam, M, Sigel, EM, Der, JP, Pittermann, J, Burge, DO, Pokorny, L, Larsson, A, Chen, T, Weststrand, S, Thomas, P, Carpenter, E, Zhang, Y, Tian, Z, Chen, L, Yan, Z, Zhu, Y, Sun, X, Wang, J, Wang, J, Stevenson, DW, Crandall-Stotler, BJ, Shaw, AJ, Deyholos, MK, Soltis, DE, Graham, SW, Windham, MD, Langdale, JA, Wong, GK-S, Mathews, S, and Pryer, KM. "Horizontal transfer of an adaptive chimeric photoreceptor from bryophytes to ferns." Proceedings of the National Academy of Sciences of the United States of America 111.18 (May 2014): 6672-6677. Full Text
  - Korall, P, and Pryer, KM. "Global biogeography of scaly tree ferns (Cyatheaceae): Evidence for Gondwanan vicariance and limited transoceanic dispersal." Journal of Biogeography 41.2 (February 1, 2014): 402-413. Full Text
  - Li, F-W, and Pryer, KM. "Crowdfunding the Azolla fern genome project: a grassroots approach." GigaScience 3 (January 2014): 16-. Full Text Open Access Copy
  - Sessa, EB, Banks, JA, Barker, MS, Der, JP, Duffy, AM, Graham, SW, Hasebe, M, Langdale, J, Li, F-W, Marchant, DB, Pryer, KM, Rothfels, CJ, Roux, SJ, Salmi, ML, Sigel, EM, Soltis, DE, Soltis, PS, Stevenson, DW, and Wolf, PG. "Between two fern genomes." GigaScience 3 (January 2014): 15-. (Review) Full Text Open Access Copy
  - Sigel, EM, Windham, MD, Smith, AR, Dyer, RJ, and Pryer, KM. "Rediscovery of Polypodium calirhiza (Polypodiaceae) in Mexico." Brittonia 66 (2014). Full Text
  - Sigel, EM, Windham, MD, Smith, AR, Dyer, RJ, and Pryer, KM. "Rediscovery of Polypodium calirhiza (Polypodiaceae) in Mexico." Brittonia 66.3 (January 1, 2014): 278-286. Full Text
  - Grusz, AL, Pryer, KM, Yatskievych, G, Huiet, RL, Gastony, GJ, and Windham, MD. "Refining the phylogeny of cheilanthoid ferns: The resurrection and recircumscription of Myriopteris (Pteridaceae) (Submitted)." Systematic Botany (2013). (Academic Article)
  - Korall, P, and Pryer, KM. "Global biogeography of scaly tree ferns (Cyatheaceae): evidence for Gondwanan vicariance and limited transoceanic dispersal (Accepted)." Journal of Biogeography 40 (2013): in press-. (Academic Article)
  - León, B, Rothfels, CJ, Arakaki, M, Young, KR, and Pryer, KM. "Revealing a cryptic fern distribution through DNA sequencing: Pityrogramma trifoliata in the western Andes of Peru (Submitted)." American Fern Journal 103 (2013). (Academic Article)
  - Rothfels, CJ, Larsson, A, Li, F-W, Sigel, EM, Huiet, L, Burge, DO, Ruhsam, M, Graham, SW, Stevenson, DW, Wong, GK-S, Korall, P, and Pryer, KM. "Transcriptome-mining for single-copy nuclear markers in ferns. (Published online)" PLoS One 8.10 (2013): e76957-. Full Text Link to Item
  - Rothfels, CR, Windham, MD, and Pryer, KM. "A plastid phylogeny of the cosmopolitan fern family Cystopteridaceae (Polypodiopsida) (Accepted)." Systematic Botany (2013). (Academic Article)
  - Beck, JB, Allison, JR, Pryer, KM, and Windham, MD. "Identifying multiple origins of polyploid taxa: a multilocus study of the hybrid cloak fern (Astrolepis integerrima; Pteridaceae)." Am J Bot 99.11 (November 2012): 1857-1865. Full Text Link to Item
  - Johnson, AK, Rothfels, CJ, Windham, MD, and Pryer, KM. "Unique expression of a sporophytic character on the gametophytes of notholaenid ferns (Pteridaceae)." Am J Bot 99.6 (June 2012): 1118-1124. Full Text Link to Item
  - Rothfels, CJ, Larsson, A, Kuo, L-Y, Korall, P, Chiou, W-L, and Pryer, KM. "Overcoming deep roots, fast rates, and short internodes to resolve the ancient rapid radiation of eupolypod II ferns." Syst Biol 61.3 (May 2012): 490-509. Full Text Link to Item
  - Li, F-W, Pryer, KM, and Windham, MD. "Gaga, a new fern genus segregated from cheilanthes (pteridaceae)." Systematic Botany 37.4 (2012): 845-860. Full Text Open Access Copy
  - Pryer, KM. "The distinguished legacy of DMB: Donald MacPhail Britton (1923 - 2012)." American Fern Journal 102 (2012): in press-. (Academic Article)
  - Pryer, KM. "The Distinguished Legacy of DMB: Donald MacPhail Britton (1923-2012)." AMERICAN FERN JOURNAL 102.4 (2012): 241-251. Full Text Link to Item
  - Rothfels, CJ, Sundue, MA, Kato, M, Larsson, A, Kuo, L-Y, Schuettpelz, E, and Pryer, KM. "A revised classification for eupolypod II ferns (Polypodiales: Polypodiopsida)." Taxon 61 (2012): 515-533. (Academic Article)
  - Rothfels, CJ, Sundue, MA, Kuo, L-Y, Larsson, A, Kato, M, Schuettpelz, E, and Pryer, KM. "A revised family-level classification for eupolypod II ferns (Polypodiidae: Polypodiales)." Taxon 61.3 (2012): 515-533.
  - Beck, JB, Windham, MD, and Pryer, KM. "Do asexual polyploid lineages lead short evolutionary lives? A case study from the fern genus Astrolepis." Evolution 65.11 (November 2011): 3217-3229. Full Text Link to Item
  - Wolf, PG, Der, JP, Duffy, AM, Davidson, JB, Grusz, AL, and Pryer, KM. "The evolution of chloroplast genes and genomes in ferns." Plant Mol Biol 76.3-5 (July 2011): 251-261. Full Text Link to Item
  - Li, F-W, Kuo, L-Y, Rothfels, CJ, Ebihara, A, Chiou, W-L, Windham, MD, and Pryer, KM. "rbcL and matK earn two thumbs up as the core DNA barcode for ferns." PLoS One 6.10 (2011): e26597-. Full Text Link to Item
  - Sigel, EM, Windham, MD, Huiet, L, Yatskievych, G, and Pryer, KM. "Species relationships and farina evolution in the cheilanthoid fern genus Argyrochosma (Pteridaceae)." Systematic Botany 36.3 (2011): 554-564. Full Text
  - Pryer, KM, Schuettpelz, E, Huiet, L, Grusz, AL, Rothfels, CJ, Avent, T, Schwartz, D, and Windham, MD. "DNA barcoding exposes a case of mistaken identity in the fern horticultural trade." Mol Ecol Resour 10.6 (November 2010): 979-985. Full Text Link to Item
  - Korall, P, Schuettpelz, E, and Pryer, KM. "Abrupt deceleration of molecular evolution linked to the origin of arborescence in ferns." Evolution 64.9 (September 2010): 2786-2792. Full Text Link to Item
  - Beck, JB, Windham, MD, Yatskievych, G, and Pryer, KM. "A diploids-first approach to species delimitation and interpreting polyploid evolution in the fern genus astrolepis (pteridaceae)." Systematic Botany 35.2 (2010): 223-234. Full Text
  - Grusz, AL, Windham, MD, and Pryer, KM. "Deciphering the origins of apomictic polyploids in the Cheilanthes yavapensis complex (Pteridaceae)." Am J Bot 96.9 (September 2009): 1636-1645. Full Text Link to Item
  - Schuettpelz, E, and Pryer, KM. "Evidence for a Cenozoic radiation of ferns in an angiosperm-dominated canopy." Proc Natl Acad Sci U S A 106.27 (July 7, 2009): 11200-11205. Full Text Link to Item
  - Pryer, KM, and Hearn, DJ. "Evolution of leaf form in marsileaceous ferns: evidence for heterochrony." Evolution 63.2 (February 2009): 498-513. Full Text Link to Item
  - Schneider, H, Smith, AR, and Pryer, KM. "Is morphology really at odds with molecules in estimating fern phylogeny?." Systematic Botany 34.3 (2009): 455-475. Full Text
  - Christenhusz, MJM, Tuomisto, H, Metzgar, JS, and Pryer, KM. "Evolutionary relationships within the Neotropical, eusporangiate fern genus Danaea (Marattiaceae)." Mol Phylogenet Evol 46.1 (January 2008): 34-48. Full Text Link to Item
  - Hennequin, S, Schuettpelz, E, Pryer, KM, Ebihara, A, and Dubuisson, J-Y. "Divergence times and the evolution of epiphytism in filmy ferns (Hymenophyllaceae) revisited." International Journal of Plant Sciences 169.9 (2008): 1278-1287. Full Text
  - Metzgar, JS, Skog, JE, Zimmer, EA, and Pryer, KM. "The paraphyly of Osmunda is confirmed by phylogenetic analyses of seven plastid loci." Systematic Botany 33.1 (2008): 31-36. Full Text
  - Nagalingum, NS, Nowak, MD, and Pryer, KM. "Assessing phylogenetic relationships in extant heterosporous ferns (Salviniales), with a focus on Pilularia and Salvinia." Botanical Journal of the Linnean Society 157.4 (2008): 673-685. Full Text
  - Rothfels, CJ, Windham, MD, Grusz, AL, Gastony, GJ, and Pryer, KM. "Toward a monophyletic Notholaena (Pteridaceae): Resolving patterns of evolutionary convergence in xeric-adapted ferns." Taxon 57.3 (2008): 712-724.
  - Schuettpelz, E, Grusz, AL, Windham, MD, and Pryer, KM. "The utility of nuclear gapCp in resolving polyploid fern origins." Systematic Botany 33.4 (2008): 621-629. Full Text
  - Haugen, P, Bhattacharya, D, Palmer, JD, Turner, S, Lewis, LA, and Pryer, KM. "Cyanobacterial ribosomal RNA genes with multiple, endonuclease-encoding group I introns. (Published online)" BMC Evol Biol 7 (September 8, 2007): 159-. Full Text Link to Item
  - Schuettpelz, E, Schneider, H, Huiet, L, Windham, MD, and Pryer, KM. "A molecular phylogeny of the fern family Pteridaceae: assessing overall relationships and the affinities of previously unsampled genera." Mol Phylogenet Evol 44.3 (September 2007): 1172-1185. Full Text Link to Item
  - Korall, P, Conant, DS, Metzgar, JS, Schneider, H, and Pryer, KM. "A molecular phylogeny of scaly tree ferns (Cyatheaceae)." Am J Bot 94.5 (May 2007): 873-886. Full Text Link to Item
  - Metzgar, JS, Schneider, H, and Pryer, KM. "Phylogeny and divergence time estimates for the fern genus Azolla (Salviniaceae)." International Journal of Plant Sciences 168.7 (2007): 1045-1053. Full Text
  - Nagalingum, NS, Schneider, H, and Pryer, KM. "Molecular phylogenetic relationships and morphological evolution in the heterosporous fern genus Marsilea." Systematic Botany 32.1 (2007): 16-25. Full Text
  - Schuettpelz, E, and Pryer, KM. "Fern phylogeny inferred from 400 leptosporangiate species and three plastid genes." Taxon 56.4 (2007): 1037-1050.
  - Korall, P, Pryer, KM, Metzgar, JS, Schneider, H, and Conant, DS. "Tree ferns: monophyletic groups and their relationships as revealed by four protein-coding plastid loci." Mol Phylogenet Evol 39.3 (June 2006): 830-845. Full Text Link to Item
  - Schuettpelz, E, and Pryer, KM. "Reconciling extreme branch length differences: decoupling time and rate through the evolutionary history of filmy ferns." Syst Biol 55.3 (June 2006): 485-502. Full Text Link to Item
  - Korall, P, Conant, DS, Schneider, H, Ueda, K, Nishida, H, and Pryer, KM. "On the phylogenetic position of Cystodium: It's not a tree fern - It's a polypod!." American Fern Journal 96.2 (2006): 45-53. Full Text
  - Nagalingum, NS, Schneider, H, and Pryer, KM. "Comparative morphology of reproductive structures in heterosporous water ferns and a reevaluation of the sporocarp." International Journal of Plant Sciences 167.4 (2006): 805-815. Full Text
  - Schuettpelz, E, Korall, P, and Pryer, KM. "Plastid atpA data provide improved support for deep relationships among ferns." Taxon 55.4 (2006): 897-906.
  - Smith, AR, Pryer, KM, Schuettpelz, E, Korall, P, Schneider, H, and Wolf, PG. "A classification for extant ferns." Taxon 55.3 (2006): 705-731.
  - Wikström, N, and Pryer, KM. "Incongruence between primary sequence data and the distribution of a mitochondrial atp1 group II intron among ferns and horsetails." Mol Phylogenet Evol 36.3 (September 2005): 484-493. Full Text Link to Item
  - Pryer, KM, Schuettpelz, E, Wolf, PG, Schneider, H, Smith, AR, and Cranfill, R. "Phylogeny and evolution of ferns (monilophytes) with a focus on the early leptosporangiate divergences." Am J Bot 91.10 (October 2004): 1582-1598. Full Text Link to Item
  - Schneider, H, Schuettpelz, E, Pryer, KM, Cranfill, R, Magallón, S, and Lupia, R. "Ferns diversified in the shadow of angiosperms." Nature 428.6982 (April 1, 2004): 553-557. Full Text Link to Item
  - Dubuisson, J-Y, Hennequin, S, Douzery, EJP, Cranfill, RB, Smith, AR, and Pryer, KM. "rbcL phylogeny of the fern genus Trichomanes (Hymenophyllaceae), with special reference to Neotropical taxa." International Journal of Plant Sciences 164.5 (2003): 753-761.
  - Marais, DLD, Smith, AR, Britton, DM, and Pryer, KM. "Phylogenetic relationships and evolution of extant horsetails, Equisetum, based on chloroplast DNA sequence data (rbcL and trnL-F)." International Journal of Plant Sciences 164.5 (2003): 737-751.
  - Pryer, KM, Schneider, H, Zimmer, EA, and Ann Banks, J. "Deciding among green plants for whole genome studies." Trends Plant Sci 7.12 (December 2002): 550-554. Link to Item
  - Schneider, H, and Pryer, KM. "Structure and function of spores in the aquatic heterosporous fern family Marsileaceae." International Journal of Plant Sciences 163.4 (2002): 485-505. Full Text
  - Pryer, KM, Smith, AR, Hunt, JS, and Dubuisson, JY. "rbcL data reveal two monophyletic groups of filmy ferns (Filicopsida: Hymenophyllaceae)." Am J Bot 88.6 (June 2001): 1118-1130. Link to Item
  - Pryer, KM, Schneider, H, Smith, AR, Cranfill, R, Wolf, PG, Hunt, JS, and Sipes, SD. "Horsetails and ferns are a monophyletic group and the closest living relatives to seed plants." Nature 409.6820 (February 1, 2001): 618-622. Full Text Link to Item
  - PRYER, KM. "Horsetails and ferns are a monophyletic group and the closest living relatives to seed plants." Nature 409 (2001): 618-622. Full Text
  - Smith, AR, Tuomisto, H, Pryer, KM, Hunt, JS, and Wolf, PG. "Metaxya lanosa, a second species in the genus and fern family Metaxyaceae." Systematic Botany 26.3 (2001): 480-486.
  - Lupia, R, Schneider, H, Moeser, GM, Pryer, KM, and Crane, PR. "Marsileaceae sporocarps and spores from the late cretaceous of Georgia, U.S.A." International Journal of Plant Sciences 161.6 (2000): 975-988. Full Text
  - Pryer, KM. "Phylogeny of Marsileaceous Ferns and Relationships of the Fossil Hydropteris pinnata Reconsidered." Int J Plant Sci 160.5 (September 1999): 931-954. Link to Item
  - Turner, S, Pryer, KM, Miao, VP, and Palmer, JD. "Investigating deep phylogenetic relationships among cyanobacteria and plastids by small subunit rRNA sequence analysis." J Eukaryot Microbiol 46.4 (July 1999): 327-338. Link to Item
  - Wolf, PG, Sipes, SD, White, MR, Martines, ML, Pryer, KM, Smith, AR, and Ueda, K. "Phylogenetic relationships of the enigmatic fern families Hymenophyllopsidaceae and Lophosoriaceae: Evidence from rbcL nucleotide sequences." Plant Systematics and Evolution 219.3-4 (1999): 263-270.
  - Hasebe, M, Wolf, PG, Pryer, KM, Ueda, K, Ito, M, Sano, R, Gastony, GJ, Yokoyama, J, Manhart, JR, Murakami, N, Crane, EH, Haufler, CH, and Hauk, WD. "Fern phylogeny based on rbcL nucleotide sequences." AMERICAN FERN JOURNAL 85.4 (1995): 134-181. Full Text Link to Item
  - Pryer, KM, Smith, AR, and Skog, JE. "Phylogenetic relationships of extant ferns based on evidence from morphology and rbcL sequences." AMERICAN FERN JOURNAL 85.4 (1995): 205-282. Full Text Link to Item
  - PRYER, KM, and HAUFLER, CH. "ISOZYMIC AND CHROMOSOMAL EVIDENCE FOR THE ALLOTETRAPLOID ORIGIN OF GYMNOCARPIUM-DRYOPTERIS (DRYOPTERIDACEAE)." SYSTEMATIC BOTANY 18.1 (1993): 150-172. Full Text Link to Item
  - PRYER, KM. "THE STATUS OF GYMNOCARPIUM-HETEROSPORUM AND G-ROBERTIANUM IN PENNSYLVANIA." AMERICAN FERN JOURNAL 82.1 (1992): 34-39. Full Text Link to Item
  - Pryer, K. "The limestone oak fern: new to Manitoba." The Blue Jay 48.4 (1990): 192-195. (Academic Article)
  - Pryer, KM, and Phillippe, LR. "A synopsis of the genus Sanicula (Apiaceae) in eastern Canada." Canadian Journal of Botany 67.3 (1989): 694-707.
  - MCNEILL, J, and PRYER, KM. "THE STATUS AND TYPIFICATION OF PHEGOPTERIS AND GYMNOCARPIUM." TAXON 34.1 (1985): 136-143. Full Text Link to Item
  - PRYER, KM, BRITTON, DM, and MCNEILL, J. "A NUMERICAL-ANALYSIS OF CHROMATOGRAPHIC PROFILES IN NORTH-AMERICAN TAXA OF THE FERN GENUS GYMNOCARPIUM." CANADIAN JOURNAL OF BOTANY-REVUE CANADIENNE DE BOTANIQUE 61.10 (1983): 2592-2602. Full Text Link to Item
  - PRYER, KM, and BRITTON, DM. "SPORE STUDIES IN THE GENUS GYMNOCARPIUM." CANADIAN JOURNAL OF BOTANY-REVUE CANADIENNE DE BOTANIQUE 61.2 (1983): 377-388. Full Text Link to Item
  - Sarvela, J, Britton, DM, and Pryer, KM. "Studies on the Gymnocarpium robertianum complex in North America." Rhodora 83 (1981): 421-431. (Academic Article)
  - Chase, R, Pryer, K, Baker, R, and Madison, D. "Responses to conspecific chemical stimuli in the terrestrial snail Achatina fulica (Pulmonata: Sigmurethra)." Behavioral Biology 22.3 (1978): 302-315.
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